Prejeto / Received: 12.11.2012 Sprejeto / Accepted: 2.12.2012 Contribution to the knowledge on the distribution of Recent free-living freshwater ostracods (Podocopida, Ostracoda, Crustacea) in Slovenia Nataša MORI & Claude MEISCH National Institute of Biology, Večna pot 111, SI-1000 Ljubljana, Slovenia; E-mail: natasa.mori@nib.si Musee national d'histoire naturelle de Luxembourg, 25, rue Munster, L-2160 Luxembourg, Luxembourg Abstract. An updated checklist of Recent free-living freshwater ostracods (Podocopida, Ostracoda, Crustacea) from Slovenia together with new records is presented. The new checklist is based on both the records extracted from the literature and the sampling in the field carried out during the last decade. The present checklist comprises 61 species. However, the eastern part of Slovenia and the surface waters are underrepresented. It is expected that the number of species will increase in the future. Key words: Ostracoda, species, fauna, freshwaters, groundwaters Izvleček. PRISPEVEK K POZNAVANJU RAZŠIRJENOSTI RECENTNIH PROSTOŽIVEČIH DVOKLOPNIKOV (PODOCOPIDA, OSTRACODA, CRUSTACEA) V CELINSKIH VODAH SLOVENIJE - Avtorja predstavljata najnovejši seznam recentnih vrst prostoživečih dvoklopnikov (Podocopida, Ostracoda, Crustacea) iz celinskih voda Slovenije ter nove najdbe za Slovenijo. Novi seznam temelji na podatkih, povzetih iz obstoječih publikacij, ter podatkih, pridobljenih med terenskim vzorčenjem v zadnjih 10 letih, ter vsebuje 61 vrst. Pričakovati je, da je število vrst veliko večje, saj primanjkuje podatkov iz vzhodne Slovenije in iz površinskih vod, ki so manj intenzivno vzorčevane kot podzemne vode. Ključne besede: Ostracoda, vrste, favna, celinske vode, podzemne vode NATURA SLOVENIAE 14(2): 5-22 Biotehniška fakulteta v Ljubljani in Nacionalni inštitut za biologijo, Ljubljana, 2012 Introduction Despite the fact that Slovenia is being recognized as a biodiversity hotspot, a preliminary list of Recent free-living freshwater ostracods (Podocopida, Ostracoda, Crustacea) from Slovenia reports only 47 species (Griffiths & Brancelj 1996a). For comparison, in Western Europe, without the Mediterranean and Dinaric regions, 158 species (Meisch 2000) and, at the global scale, 2090 species belonging to about 209 genera are known at present (Martens & Savatenalinton 2011). The first known record of ostracods for Slovenia is from the cave Podpeška jama at Dobrepolje near Velike Lašče, where stygobiotic species »similar to Cyclocypris ovum but flatter« was mentioned (Joseph 1882). The next report is from Postojna, where two species were collected from the larger dripping pool in the cave Črna jama (Schmeil 1893). The species were named Typhiocypris schmeiiiand Cyprispellucida by G.W. Müller, but later never described. In 1920, Paris reported on Typhlocypris eremita (Veydovsky, 1880), again from the cave Podpeška jama (Paris 1920). In the 1930's, Walter Klie described four new species from the caves Podpeška jama (Candona aemonae Klie, 1935; Cypria stygia Klie, 1935), Postojnska jama (Candona trigonella Klie, 1931) and Krška jama (Candona cavicola Klie, 1935) and an additional one from the Krka River near Novo mesto (Leptocythere fluviatilis Klie, 1939) (Klie 1931, 1935, 1939). Moreover, W. Klie received from S. Karaman the material from the cave Luknja near Novo mesto and identified it as Notodromaspersicae Gurney (Klie 1938). At the same time, H.J. Stammer referred in his work on the fauna from the Timavo River to several species collected in Submediterranean Slovenia (Stammer 1932). More recently, T. Petkovski published scattered information on the occurrence of ostracods in Slovenia within his reports on Yugoslav fauna (Petkovski 1958, 1959, 1960a, b, 1964, 1969). Two new species were described by D. Danielopol in 1978 (Pseudocandona pretneri Danielopol, 1978) and T. Petkovski & C. Meisch in 1994 (Cypria bicoiorPetkovski & Meisch, 1994). The first species was collected from the spring Rak near Cerknica (Danielopol 1982) and the second one from the spring Ižica near Ljubljana (Petkovski & Meisch 1994). At the end of the last century, H. Griffiths carried out a sampling campaign of mostly Alpine lakes and co-authored the preliminary checklist for Slovenia (Griffiths & Brancelj 1996a) as well as a paper about the palaeo-biogeography of Candona bimucronata in the Balkans, including Slovenia (Griffiths & Brancelj 1996b). In 2002, an extensive field sampling was carried out in the karst and alluvial aquifers south of Ljubljana with the aim to assess the groundwater biodiversity of the area and compare it to other European regions (EU PASCALIS project, Dole-Olivier et al. 2009, Mori et al. 2011a). On that occasion, the present authors started to collaborate on the taxonomy and distribution of the freshwater ostracods in Slovenia. After that, ostracods were continuously collected and identified during several field samplings carried out by the National Institute of Biology (Mori & Brancelj 2011, Mori et al. 2011b, 2012). The present paper summarizes the history of ostracod research in Slovenia and presents an overview of all the species recorded in this country until the present. Additionally, it provides a list of unpublished new records for Slovenia, with information on their geographical location and habitat type. The species list includes the species that were previously reported in the literature and records that were obtained through field sampling by the first author during the last 10 years. It provides general information on geographical distribution, the habitats where species were collected and lists all the publications in which the species were reported. This work provides an updated information on the occurrence of this neglected crustacean group in Slovenia and demonstrates, as it was shown by many other specialists, that Slovenia is a biodiversity hotspot (Sket 1999) also as far as ostracods are concerned. Material in methods Only Recent free-living freshwater ostracods were considered in this work. First, a careful examination of the existing original literature reporting on the occurrence of ostracods in Slovenia during the last two centuries was carried out. Based on this literature, a new species list was composed in order to update the nomenclature and to avoid possible errors from the preliminary check list published by Griffiths & Brancelj (1996a). Next, the data obtained from the first author's field samplings, starting in 2002, are included. Hence, the present species list contains all the species reported in the literature published since 1920 (Brancelj et al. 1995, Danielopol 1982, Dole-Olivier et al. 2009, Griffiths & Brancelj 1996a, b, Klie 1931, 1935, 1938, 1939, Mori & Brancelj 2011, Mori et al. 2011a, b, 2012, Paris 1920, Petkovski 1959, 1960a, b, 1964, 1969, Petkovski & Meisch 1994, Sket & Velkovrh 1981), as well as so far unpublished species records of the first author from 2002 to the present. The species are listed according to the classification proposed by Martens & Savatenalinton (2011). Only the most important synonyms are listed, more of them are available in Martens & Savatenalinton (2011). The species with nomen nudum are discarded from the list to avoid confusion (Joseph 1882, Schmeil 1893). The list of locations for the species collected by the first author from 2002 onwards is provided. That list contains the geographical name of the corresponding location, the habitat type, Gauss-Krüger coordinates, altitude, date of collecting and the name of the persons who collected the sample and identified the species. The localities selected during the EU PASCALIS project were sampled by several different field workers from NIB and Biotechnical Faculty in Ljubljana and their names are not specified. In the species list, a valid species name, the general information on geographical distribution based on the phytogeographic division of Slovenia (AL - Alpine, SA - Subalpine, SM - Submediterranean, DN - Dinaric, SD - Subdinaric, SP - Subpannonian) (Wraber 1969) and habitats types where the species was collected are provided. Additionally, for each species, the publications where the species was reported until present are cited. For each genus, the type species is marked by asterisk (*). List of species Class Ostracoda Latreille, 1806 Subclass Podocopa G.W. Müller, 1894 Order Podocopida G.O. Sars, 1866 Superfamily Cypridoidea Baird, 1845 Family Cyprididae Baird, 1845 Subfamily Cypricercinae McKenzie, 1971 *Bradleycypris obliqua (Brady, 1868) [Cypris obliqua (Brady, 1868)] AL, DN - lake littoral - Griffiths & Brancelj 1996a Subfamily Cypridopsinae Kaufmann, 1900 Cavernocypris subterranea (Wolf, 1920) [Cypridopsis subterraneaWolf, 1920] AL, DN, SM - spring benthos and drift, lake littoral, caves - Brancelj et al. 1995, Griffiths & Brancelj 1996a, Mori et al. 2011b *Cypridopsis vidua (O.F. Müller, 1776) [Cypris vidua (O.F. Müller, 1776)] AL, DN, SD, SM - lake littoral, swamp, river benthos, hyporheic zone, caves - Klie 1931, Stammer 1932, Griffiths & Brancelj 1996a Potamocypris fallax Fox, 1967 AL - river and spring benthos - Griffiths & Brancelj 1996a *Potamocypris fulva (Brady, 1868) [Bairdia f/lva Brady, 1868] AL, DN - spring benthos and drift, hyporheic zone - Mori et al. 2011b Potamocypris pallida Alm, 1914 AL - spring benthos - Mori et al. 2011b Potamocypris simiiisG.W. Müller, 1912 AL - spring benthos, swamp - Griffiths & Brancelj 1996a Potamocypris smaragdina (Vavra, 1891) [CypridopsissmaragdinaVavra, 1891] AL - swamp - Griffiths & Brancelj 1996a Potamocypris variegata (Brady & Norman, 1889) [Cypridopsis variegata Brady & Norman, 1889] SD - river benthos Potamocypris villosa (Jurine, 1820) [Monoculus villosa Jurine, 1820] SM - cave, spring - Stammer 1932, Griffiths & Brancelj 1996a Potamocypris zschokkei (Kaufmann, 1900) [Paracypridopsis zschokkei Kaufmann, 1900] AL, DN - spring benthos and hyporheic zone of the headwaters - Brancelj et al. 1995, Griffiths & Brancelj 1996a, b, Dole-Olivier et al. 2009, Mori et al. 2011b Subfamily Cyprinotinae Bronshtein, 1947 *Heterocypris incongruens (Ramdohr, 1808) [Cypris incongruens Ramdohr, 1808] AL - ponds, lake littoral - Griffiths & Brancelj 1996a Heterocypris reptans (Kaufmann, 1900) [Microcypris reptans Kaufmann, 1900] DN, SM - spring benthos - Stammer 1932, Petkovski 1964 Subfamily Dolerocypridinae Triebel, 1961 Doierocyprissinensis G.O. Sars 1903 SA/SD - pit overgrown by dense vegetation - Petkovski 1960a Subfamily Eucypridinae Bronshtein, 1947 Eucyprispigra (Fischer, 1851) [Cyprispigra Fischer, 1851] AL, DN - benthos and hyporheic zone - Mori et al. 2011b *Eucypris virens (Jurine, 1820) [Monocu/us virens Jurine, 1820] DN - lake littoral - Griffiths & Brancelj 1996a Subfamily Herpetocypridinae Kaufmann, 1900 Psychrodromus fontina/is (Wolf, 1920) [Iyodromus fontinaiisWolf, 1920] AL - spring benthos, river benthos - Griffiths & Brancelj 1996a, Mori et al. 2011b *Psychrodromus olivaceus (Brady & Norman, 1889) [Erpetocypris olivacea Brady & Norman, 1889] AL - river benthos - Griffiths & Brancelj 1996a Subfamily Scottiinae Bronshtein, 1947 *Scottia pseudobrowniana Kempf, 1971 DN - spring benthos - Dole-Olivier et al. 2009 Family Candonidae Kaufmann, 1900 Subfamily Candoninae Kaufmann, 1900 Tribe Candonini Kaufmann, 1900 Candona bimucronata Klie, 1937 AL - lake littoral - Griffiths & Brancelj 1996a, b *Candona candida(O.F. Müller, 1776) [CypriscandidaO.F. Müller, 1776] AL, SA, DN, SD, SM - lake littoral, river benthos and hyporheic zone, spring benthos, caves - Stammer 1932, Sket & Velkovrh 1981, Brancelj et al. 1995, Griffiths & Brancelj 1996a, b, Mori & Brancelj 2011, Mori et al. 2012 Candona iindneriG.O. Petkovski, 1969 DN, SM - spring and river benthos - Petkovski 1969, Griffiths & Brancelj 1996a Candona negiecta Sars, 1887 AL, SD - lake littoral, spring benthos, hyporheic zone, moss - Brancelj et al. 1995, Griffiths & Brancelj 1996a *Cryptocandona vavrai Kaufman, 1900 AL, SA, DN - spring benthos, hyporheic zone - Mori et al. 2011b Fabaeformiscandona aemonae (Klie, 1935) [Candona aemonaeKlie, 1935] AL, SA, DN, SD - spring benthos, hyporheic zone, caves - Klie 1935, Dole-Olivier et al. 2009, Mori et al. 2011a, 2012 Fabaeformiscandona breuiii (Paris, 1920) [CandonabreuHi Paris, 1920] AL, SA, DN - spring benthos, hyporheic zone - Mori & Brancelj 2011, Mori et al. 2011b Fabaeformiscandona brevicornis (Klie, 1925) [Candona brevicornis Klie, 1925] DN - spring benthos, hyporheic zone Fabaeformiscandona brisiaca (Klie, 1938) [Candona brisiaca Klie, 1938] AL - spring drift - Mori et al. 2011b Fabaeformiscandona latens (Klie, 1940) [Candona iatens Klie, 1940] AL- spring benthos - Mori et al. 2011b. Mixtacandona chappuisi(Klie, 1943) [CandonachappuisiKlie, 1943] DN - spring benthos, hyporheic zone - Dole-Olivier et al. 2009 Mixtacandona iattingerae Rogulj & Danielopol 1993 DN - spring benthos - Dole-Olivier et al. 2009 Mixtacandona laisi(K\ie, 1938) [Candona laisiKlie, 1938] Syn.: Candona stammeri Klie, 1938 DN - spring benthos, hyporheic zone - Dole-Olivier et al. 2009, Mori et al. 2011b *Nannocandona faba Eckman, 1914 AL - lake littoral - Brancelj et al. 1995, Griffiths & Brancelj 1996a. Pseudocandona albicans (Brady, 1864) [Candona albicans Brady, 1864] AL, SA, DN, SD, SM - caves, spring benthos, hyporheic zone - Griffiths & Brancelj 1996a Pseudocandona lobipes(Hartwig, 1900) [Candona lobipes Hartwig, 1900] SD - Sphagnum moss Pseudocandona pratensis (Hartwig, 1901) [Candona pratensis Hartwig, 1901] DN - lake littoral Typhlocypris cavicoaa (Klie, 1935) [Candona cavico/a Klie, 1935] DN - caves, spring benthos, hyporheic zone - Klie 1935, Dole-Olivier et al. 2009 Typhlocypris eremita (Veydovskyy, 1880) [Cypris eremita Veydovsky, 1880] DN - cave - Paris 1920, Dole-Olivier et al. 2009 Typhlocypris pretneri Danielopol, 1978 DN - spring drift - Danielopol 1978, Danielopol 1982, Dole-Olivier et al. 2009 Typhlocypris trigonella (Klie, 1931) [Candona trigonella Klie, 1931] DN - pond in a cave - Klie 1931 Tribe Candonopsini Karanovic, 2004 Candonopsis scourfieldi Brady, 1910 SA, DN - spring benthos and hyporheic zone Subfamily Cyclocypridinae Kaufmann, 1900 *Cyclocypris globosa (Sars, 1863) [Cypris globosa Sars, 1863] SD - Sphagnum moss Cyclocyprislaevis(O.F. Müller, 1776) [CyprislaevisO.F. Müller, 1776] AL, DN - lake littoral and spring benthos - Griffiths & Brancelj 1996a Cyclocypris ovum (Jurine, 1820) [Monoculusovum Jurine, 1820] AL, SA - lake littoral, river benthos, Brancelj et al. 1995, Griffiths & Brancelj 1996a, b Cypria bicolor Petkovski & Meisch, 1994 DN - spring benthos - Petkovski & Meisch 1994 Cypria cavernae Wagenleitner, 1990 DN - spring benthos - Dole-Olivier et al. 2009. Cypria exsculpta Fischer, 1855 [Cypris exsculpta Fischer, 1855] AL - lake littoral - Griffiths & Brancelj 1996a Cypria lacustris Lilljeborg, 1890 SA, DN, SA - well, spring benthos, hyporheic zone - Petkovski 1960b Cypria ophtalmica (Jurine, 1820) [Monoculus ophtalmica Jurine, 1820] AL, SA, DN, SD, SM - lake littoral, spring benthos, hyporheic zone, caves - Klie 1931, 1939, Stammer 1932, Klie 1939, Brancelj et al. 1995, Griffiths & Brancelj 1996a, b Cypria reptans Brohnstein, 1928 AL, DN - caves, spring benthos, hyporheic zone - Klie 1935, Petkovski & Meisch 1994, Griffiths & Brancelj 1996a Cypria sketi Petkovski, 1976 DN - spring benthos, hyporheic zone, pond Family Ilyocyprididae Kaufmann, 1900 Subfamily Ilyocypridinae Kaufmann, 1900 Ityocypris bradyi G.O.Sars 1890 SA, DN, SD - Petkovski 1958, Griffiths & Brancelj 1996a. *Ityocyprisgibba (Ramdohr, 1808) [Cyprisgibba Ramdohr, 1808] AL, SA - Petkovski 1958, Griffiths & Brancelj 1996a Ityocypris inermis Kaufmann, 1900 SM - spring benthos - Griffiths & Brancelj 1996a Family Notodromadidae Kaufmann, 1900 Subfamily Notodromadinae Kaufmann, 1900 *Notodromasmonacha(O.F. Müller, 1776) [CyprismonachaO.F. Müller, 1776] AL - pond - Petkovski 1959 Notodromas persica Gurney, 1921 DN - cave - Klie 1938 Superfamily Darwinuloidea Brady & Roberston, 1885 Family Darwinulidae Brady & Robertson, 1885 *Darwinula stevensoni (Brady & Robertson, 1870) [Polycheles stevensoni Brady & Norman, 1889] AL - lake littoral - Griffiths & Brancelj 1996a Superfamily Cytheroidea Baird, 1850 Family Cytherideidae Baird, 1850 Subfamily Cytherideinae Sars, 1925 *Cytherissa lacustris(Sars, 1863) [Cythere lacustrisSars, 1863] AL - lake littoral - Griffiths & Brancelj 1996a, b Family Leptocytheridae Sars, 1925 Leptocythiere flu viatilis Klie, 1939 DN - river benthos - Klie 1939 Family Limnocytheridae Klie, 1938 Subfamily Limnocytherini Klie, 1938 *Limnocythere inopinata (Baird, 1843) [Cythere inopinata Baird, 1843] AL; swamp, lake littoral - Griffiths & Brancelj 1996a. Limnocytherina sanctipatricii(Brady & Robertson, 1869) [Limnocythere sanctipatricii Brady & Robertson, 1869] AL - lake littoral - Griffiths & Brancelj 1996a, b New records for Slovenia 1. Jama ob Savi, Brestanica, cave, 5093457 5531401, 169 m a. s. l., 17.1.05, leg. A. Kapla, det. N. Mori: Typhiocypris cavicola 2. Škocjanske jame, Divača, cave, 5058470 5421620, 320 m a. s. l., 1.2.05, leg. A. Kapla, det. C. Meisch: Typhiocypris cavicola 3. Škocjanske jame, Divača, cave, 5058470 5421620, 320 m a. s. l., 17.2.05, leg. A. Kapla, det. N. Mori: Cavernocypris subterranea 4. Turjeva jama, Kobarid, cave, 5123233 5385050, 253 m a. s. l., 20.3.05, leg. A. Kapla, det. N. Mori: Fabaeformiscandona aemonae 5. Velika Pasica, Ig, cave, 5086180 5461052, 700 m a. s. l., 26.9.07, leg. A. Brancelj, det. N. Mori: Pseudocandona albicans 6. Slabetova jama, Vrhnika, cave, 5094862 5440268, 436 m a. s. l., 1.9.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandona aemonae 7. Lipnik spring, Bled, benthos, 5138308 5425554, 650 m a. s. l., 10.6.09, leg. B. Oz, det. N. Mori: Cavernocypris subterranea 8. Lipnik spring, Bled, benthos, 5138308 5425554, 650 m a. s. l., 4.11.09, leg. B. Oz, det. N. Mori: Fabaeformiscandona breuili 9. Lipnik spring, Bled, benthos, 5138308 5425554, 650 m a. s. l., 26.4.12, leg. M. Opalički, det. N. Mori: Psychrodromus fontinalis 10. spring at Breg pri Borovnici, Borovnica, benthos, 5088350 5451050, 300 m a. s. l., 19.8.02, Pascalis 2002, det. C. Meisch: Cypriasketi Mixtacandona laisi, Pseudocandona albicans, Typhiocypris cavicola 11. spring at Šivčev grič, Borovnica, benthos, 5091450 5452950, 290 m a. s. l., 21.8.02, Pascalis 2002, det. C. Meisch: Cypria sketi 12. spring at Podkamnik, Borovnica, benthos, 5091250 5454700, 295 m a. s. l., 21.8.02, Pascalis 2002, det. C. Meisch: Cypria reptans, Heterocyprisreptans, Pseudocandona albicans, Typhiocypris cavicoaa 13. spring at Preserje, Borovnica, benthos, 5091625 5454825, 295 m a. s. l., 21.8.02, Pascalis 2002, det. C. Meisch: Typhiocypris cavicoaa 14. spring at Ribič, Borovnica, benthos, 5088525 5448900, 300 m a. s. l., 29.8.02, Pascalis 2002, det. C. Meisch: Cypria reptans, Fabaeformiscandona aemonae 15. spring at Dol pri Borovnici, Borovnica, benthos, 5087500 5449875, 300 m a. s. l., 29.8.02, Pascalis 2002, det. C. Meisch: Cypria reptans, Hyocyprisbradyi, Pseudocandona albicans, Typhiocypris cavicoaa 16. spring at Ohonica, Borovnica, benthos, 5084325 5451775, 340 m a. s. l., 29.8.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi 17. spring at Malence, Borovnica, benthos, 5085450 5450375, 450 m a. s. l., 6.9.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi, Scottia pseudobrowniana 18. spring Veliko Bojišče, Borovnica, benthos, 5084775 5450825, 500 m a. s. l., 6.9.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandona brevicornis, Fabaeformiscandona breuili 19. spring pod Sv. Jožefom, Borovnica, benthos, 5090850 5454650, 295 m a. s. l., 16.9.02, Pascalis 2002, det. C. Meisch: Cypria reptans 20. spring at Ponikve, Borovnica, spring, 5495550 5455450, 300 m a. s. l., 10.12.2002, Pascalis 2002, det. C. Meisch: Typhlocypris cavicola 21. spring Kotliči, Cerknica, benthos, 5072085 5445511, 600 m a. s. l., 1.7.11, leg. N. Mori, det. N. Mori: Typhlocypris cavicola 22. spring Rak, Cerknica, benthos, 5072479 5446769, 640 m a. s. l., 28.7.11, leg. N. Mori, det. N. Mori: Typhlocypris cavicola 23. Dobličko jezero, Črnomelj, benthos, 5045288 5511640, 200 m a. s. l., 12.7.05, leg. A. Kapla, det. N. Mori: Cypria sketi 24. spring Mrzlica, Ig, benthos, 5085650 5460000, 760 m a. s. l., 26.8.02, Pascalis 2002, det. C. Meisch: Potamocypris fuiva 25. spring under Mišnice, Ig, benthos, 5084775 5459675, 800 m a. s. l., 26.8.02, Pascalis 2002, det. C. Meisch: Potamocypris fuiva 26. spring Gabrovica, Ig, benthos, 5084500 5462775, 680 m a. s. l., 29.8.02, Pascalis 2002, det. C. Meisch: Cypria reptans, Potamocypris fuiva 27. spring Korito, Ig, benthos, 5078800 5462275, 690 m a. s. l., 3.9.02, Pascalis 2002, det. C. Meisch: Candona neglecta, Cypria reptans 28. spring at Ižanka, Ig, benthos, 5092100 5463650, 290 m a. s. l., 8.9.02, Pascalis 2002, det. C. Meisch: Cyclocypris laevis, Cypria reptans 29. spring in Draga, Ig, benthos, 5087750 5465500, 320 m a. s. l., 19.9.02, Pascalis 2002, det. C. Meisch: Cypria reptans, Mixtacandona laisi 30. spring at Krška jama cave, Ivančna Gorica, benthos, 5082866 5482680, 272 m a. s. l., 28.7.11, leg. N. Mori, det. N. Mori: Typhlocypris cavicola 31. spring in the Vrata valley, Jesenice, benthos, 5144888 5416086, 720 m a. s. l., 9.7.09, leg. B. Oz, det. N. Mori: Cavernocypris subterranea, Potamocypris fulva, Psychrodromus fontinalis 32. Završnica spring, Jesenice, benthos, 5141732 5438568, 943 m a. s. l., 16.6.09, leg. B. Oz, det. N. Mori: Cavernocypris subterranea 33. Završnica spring, Jesenice, drift, 5141732 5438568, 943 m a. s. l., 20.10.10, leg. N. Mori, det. N. Mori: Psychrodromus fontinalis 34. Javorniški potok spring, Jesenice, benthos, 5146689 5432032, 1108 m a. s. l., 16.6.09, leg. B. Oz, det. N. Mori: Psychrodromus fontinalis 35. Javorniški potok spring, Jesenice, benthos, 5146689 5432032, 1108 m a. s. l., 4.11.09, leg. B. Oz, det. N. Mori: Cavernocypris subterranea 36. Kamniška Bistrica spring, Kamnik, benthos, drift, 5131603 5468851, 600 m a. s. l., 4.6.09, leg. B. Oz, det. N. Mori: Cavernocypris subterranea, Mixtacandona laisi 37. Črna spring, Luče, benthos, 5140596 5472340, 740 m a. s. l., 1.7.09, leg. B. Oz, det. N. Mori: Cavernocypris subterranea 38. Pavličevo sedlo spring, Luče, drift, 5141860 5469638, 1232 m a. s. l., 1.7.09, leg. N. Mori, det. N. Mori: Fabaeformiscandona brisiaca, Psychrodromus fontinalis, Fabaeformiscandona latens, Cavernocypris subterranea, Cryptocandona vavrai, Potamocypris zschokkei 39. Presušnik spring, Mojstrana, benthos, 5148330 5422931, 1220 m a. s. l., 16.6.09, leg. B. Oz, det. N. Mori: Eucypris pigra 40. Presušnik spring, Mojstrana, benthos, 5148330 5422931, 1220 m a. s. l., 8.9.09, leg. B. Oz, det. N. Mori: Candona negiecta 41. Presušnik spring, Mojstrana, drift, 5148330 5422931, 1220 m a. s. l., 2.9.10, leg. N. Mori, det. N. Mori: Fabaeformiscandonabreui, Potamocypristuiva 42. Rogovilc spring, Mozirje, drift, 5139436 5477797, 669 m a. s. l., 2.7.09, leg. N. Mori, det. N. Mori: Psychrodromus fontinaiis, Potamocypriszschokkei 43. Rogovilc, Mozirje, drift, 5139436 5477797, 669 m a. s. l., 18.6.10, leg. N. Mori, det. N. Mori: Fabaeformiscandona breuHi, Cavernocypris subterranea, Potamocypris fallax 44. spring at Resnik, Podkraj, benthos, 5092750 5459750, 290 m a. s. l., 19.9.02, Pascalis 2002, det. C. Meisch: Cyclocyprislaevis, Cypriareptans 45. spring at Koprivec, Podlipa, benthos, 5095925 5448250, 300 m a. s. l., 25.10.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandona aemonae, Ityocypris bradyi 46. spring at Podpeč, Podpeč, benthos, 5092100 5455200, 295 m a. s. l., 21.8.02, Pascalis 2002, det. C. Meisch: Cypria sketi Ityocyprisbrady, Mixtacandona iattingerae, Mixtacandona iaisi, Typhiocypris cavicoia 47. spring at Podkraj, Podpeč, benthos, 5092134 5459145, 300 m a. s. l., 30.8.02, Pascalis 2002, det. C. Meisch: Cypria reptans, Typhiocypris cavicoia 48. spring at Podkraj, Podpeč, benthos, drift, 5092134 5459145, 300 m a. s. l., 2.3.12, leg. B. Mežnarič, det. N. Mori: Cypria reptans, Mixtacandona chappuisi, Typhiocypris cavicoia 49. spring at Podpeško jezero, Podpeč, benthos, 5091300 5456825, 290 m a. s. l., 30.8.02, Pascalis 2002, det. C. Meisch: Typhiocypris cavicoia, Cypria reptans, Typhiocypris cavicoia 50. spring at pond, Podpeč, benthos, 5091315 5456811, 300 m a. s. l., 6.7.12, leg. B. Mežnarič, det. N. Mori: Cypria reptans, Heterocyprisreptans 51. spring at Jezero, Podpeč, benthos, 5092050 5456125, 300 m a. s. l., 19.9.02, Pascalis 2002, det. C. Meisch: Cyclocypris iaevis, Cypria reptans, Fabaeformiscandona aemonae, Typhiocypris cavicoia 52. spring at Virje, Podpeč, benthos, 5092384 5457464, 300 m a. s. l., 25.10.02, Pascalis 2002, det. C. Meisch: Typhiocypris cavicoia 53. spring at Virje, Podpeč, drift, 300 m a. s. l., 6.7.12, leg. B. Mežnarič, det. N. Mori: Typhiocypris cavicoia 54. spring Vogli, Strahomer, benthos, 5088338 5460613, 310 m a. s. l., 30.7.02, Pascalis 2002, det. C. Meisch: Cypria reptans, Mixtacandona chappuisi 55. spring at Strahomer pond, Strahomer, benthos, 5087950 5460725, 320 m a. s. l., 25.9.02, Pascalis 2002, det. C. Meisch: Typhiocypris cavicoia 56. spring Močilo, Škrilje, benthos, 5084800 5465875, 590 m a. s. l., 29.8.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandona breuiii 57. springs at Mošenik, Tržič, drift, 5141621 5444173, 807 m a. s. l., 16.6.09, leg. N. Mori, det. N. Mori: Cavernocypris subterranea 58. springs at Mošenik, Tržič, benthos, 5141621 5444173, 807 m a. s. l., 16.6.09, leg. B. Oz, det. N. Mori: Potamocyprispallida, Potamocypriszschokkei, Psychrrodromus fontinaiis 59. spring at Ogrin, Velika Ligojna, benthos, 5094500 5445725, 295 m a. s. l., 30.8.02, Pascalis 2002, det. C. Meisch: Candonopsisscourfeldi, Pseudocandona albicans 60. spring Moravc, Velika Ligojna, benthos, 5095000 5444725, 320 m a. s. l., 30.8.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandona aemonae 61. Malo okence, Vrhnika, benthos, 5089750 5445775, 300 m a. s. l., 30.8.02, Pascalis 2002, det. C. Meisch: Cypria cavernae, Cypria lacustris, Typhlocypris cavicola, Typhlocypris eremtta 62. Veliki Močilnik, Vrhnika, benthos, 5090225 5445500, 300 m a. s. l., 30.8.02, Pascalis 2002, det. C. Meisch: Mixtacandona laisi, Typhlocypris cavicola 63. Korita on Planja, Vrhnika, benthos, 5092300 5442850, 500 m a. s. l., 9.9.02, Pascalis 2002, det. C. Meisch: Cypria reptans 64. spring in Kobilja dolina, Vrhnika, benthos, 5096150 5447875, 320 m a. s. l., 16.9.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi 65. spring at Velika Ligojna, Vrhnika, benthos, 5095150 5446300, 355 m a. s. l., 16.9.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandona breuiii, Fabaeformiscandona brevicornis, Mixtacandona chappuisi 66. spring at Mala Ligojna, Vrhnika, benthos, 5095550 5441825, 350 m a. s. l., 25.7.02, Pascalis 2002, det. C. Meisch: Cryptocandona vavrai 67. spring at Želimlje, Želimlje, benthos, 5086200 5467375, 320 m a. s. l., 30.7.02, Pascalis 2002, det. C. Meisch: Candonopsis scourfieldi 68. spring at Podreber, Želimlje, benthos, 5086550 5467650, 310 m a. s. l., 30.7.02, Pascalis 2002, det. C. Meisch: Cyclocyprislaevis, Mixtacandona chappuisi, Typhlocypris cavicola 69. spring at Turjak castle, Želimlje, benthos, 5081650 5469475, 355 m a. s. l., 19.9.02, Pascalis 2002, det. C. Meisch: Cypria reptans 70. Dremavščica spring, Želimlje, drift, 5089275 5466900, 295 m a. s. l., 30.11.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi, Typhlocypris cavicola 71. Sava, Ljubljana, HZ, 5104638 5468541, 278 m a. s. l., 27.12.04, leg. G. Bračko, det. N. Mori: Candona candida 72. Sava, Ljubljana, HZ, 5104402 5465197, 278 m a. s. l., 27.12.04, leg. G. Bračko, det. N. Mori: Typhlocypris cavicola 73. Sava, Ljubljana, HZ, 5108049 5459811, 296 m a. s. l., 13.7.07, leg. N. Mori, det. N. Mori: Candona candida 74. Sava, Ljubljana, HZ, 5108049 5459811, 296 m a. s. l., 13.7.07, leg. N. Mori, det. N. Mori: Fabaeformiscandona aemonae 75. Sava, Ljubljana, HZ, 5108079 5459762, 291 m a. s. l., 20.8.08, leg. U. Žibrat, det. N. Mori: Pseudocandona albicans 76. Sava, Naklo, HZ, 5125566 5445065, 366 m a. s. l., 20.8.08, leg. U. Žibrat, det. N. Mori: Pseudocandona albicans 77. Sava, Radovljica, HZ, 5132549 5436600, 406 m a. s. l., 20.8.08, leg. U. Zibrat, det. N. Mori: Fabaeformiscandona breuHi 78. Sava, Jevnica, HZ, 5104727 5479620, 251 m a. s. l., 21.8.08, leg. U. Zibrat, det. N. Mori: Cypria reptans 79. Želimeljščica, Želimlje, HZ, 5080873 5469709, 370 m a. s. l., 8.5.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandona breuHi 80. Želimeljščica, Želimlje, HZ, 5082311 5469014, 345 m a. s. l., 8.5.02, Pascalis 2002, det. C. Meisch: Cypria reptans, Potamocypris fulva, Pseudocandona albicans, Fabaeformiscandona breuiii 81. Želimeljščica, Želimlje, HZ, 5083125 5468825, 336 m a. s. l., 14.5.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi, Pseudocandona albicans 82. Želimeljščica, Želimlje, HZ, 5083125 5468875, 336 m a. s. l., 14.5.02, Pascalis 2002, det. C. Meisch: Cypria reptans 83. Želimeljščica, Želimlje, HZ, 5084000 5468350, 332 m a. s. l., 16.5.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi 84. Želimeljščica, Želimlje, HZ, 5084300 5468200, 331 m a. s. l., 16.5.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandonabreuüi, Mixtacandona chappuisi, Pseudocandona albicans, Typhlocypris cavicola 85. Želimeljščica, Želimlje, HZ, 5085050 5467950, 320 m a. s. l., 16.5.02, Pascalis 2002, det. C. Meisch: Cypria reptans, Mixtacandona chappuisi, Pseudocandona albicans 86. Želimeljščica, Želimlje, HZ, 5086450 5467650, 309 m a. s. l., 16.5.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi, Fabaeformiscandona breuHi, Pseudocandona albicans, Typhlocypris cavicola 87. Želimeljščica, Želimlje, HZ, 5087450 5466850, 301 m a. s. l., 17.5.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandona breuHi, Fabaeformiscandona brevicornis, Pseudocandona albicans, Mixtacandona chappuisi 88. Želimeljščica, Želimlje, HZ, 5088450 5466500, 297 m a. s. l., 30.5.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi, Pseudocandona albicans 89. Želimeljščica, Želimlje, HZ, 5088998 5466383, 295 m a. s. l., 25.7.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi, Pseudocandona albicans 90. Iška, Iška vas, HZ, 5084300 5461000, 375 m a. s. l., 31.5.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandona aemonae, Fabaeformiscandona breuHi, Potamocypris zschokkei 91. Iška, Iška vas, HZ, 5084750 5461050, 363 m a. s. l., 31.5.02, Pascalis 2002, det. C. Meisch: Pseudocandona albicans 92. Iška, Iška vas, HZ, 5085250 5461425, 359 m a. s. l., 31.5.02, Pascalis 2002, det. C. Meisch: Pseudocandona albicans 93. Iška, Iška vas, HZ, 5086690 5462300, 342 m a. s. l., 12.6.02, Pascalis 2002, det. C. Meisch: Pseudocandona albicans, Cypria sketi, Candona candida, Fabaeformiscandona breuHi, Fabaeformiscandona brevicornis 94. Iška, Iška vas, HZ, 5088113 5462847, 327 m a. s. l., 12.6.02, Pascalis 2002, det. C. Meisch: Pseudocandona albicans 95. Iška, Iška vas, HZ, 5088501 5462130, 320 m a. s. l., 13.6.02, Pascalis 2002, det. C. Meisch: Typhiocypris cavicola 96. Iška, Iška vas, HZ, 5088641 5461752, 314 m a. s. l., 13.6.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi, Typhiocypris cavicola 97. Iška, Iška vas, HZ, 5088586 5460885, 310 m a. s. l., 13.6.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi, Typhiocypris cavicola 98. Iška, Iška vas, HZ, 5088586 5460885, 310 m a. s. l., 13.6.02, Pascalis 2002, det. C. Meisch: Eucyprispigra, Mktacandona chappuis.i, Pseudocandona albicans, Typhiocypris cavicola 99. Iška, Iška vas, HZ, 5090419 5460469, 302 m a. s. l., 18.6.02, Pascalis 2002, det. C. Meisch: Typhiocypris cavicola, Pseudocandona albicans 100. Iška, Iška vas, HZ, 5091822 5460057, 295 m a. s. l., 18.6.02, Pascalis 2002, det. C. Meisch: Typhiocypris cavicola 101. Iška, Iška vas, HZ, 5091822 5460057, 295 m a. s. l., 18.6.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi, Typhiocypris cavicola 102. Borovniščica, Borovnica, HZ, 5083142 5451708, 340 m a. s. l., 19.6.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandona aemonae, Fabaeformiscandona breuiH, Typhiocypris cavicola, Mixtacandona chappuisi, Cypria reptans, Mixtacandona laisi 103. Borovniščica, Borovnica, HZ, 5083751 5452208, 320 m a. s. l., 19.6.02, Pascalis 2002, det. C. Meisch: Cryptocandona vavra, Mixtacandona iais, Typhiocypris cavicola, Cypria reptans, Cypridopsis vidua, Fabaeformiscandona aemonae 104. Borovniščica, Borovnica, HZ, 5084334 5452301, 310 m a. s. l., 19.6.02, Pascalis 2002, det. C. Meisch: Mixtacandona laisi 105. Borovniščica, Borovnica, HZ, 5082674 5453088, 340 m a. s. l., 20.6.02, Pascalis 2002, det. C. Meisch: Typhiocypris cavicola 106. Borovniščica, Borovnica, HZ, 5084779 5452057, 309 m a. s. l., 20.6.02, Pascalis 2002, det. C. Meisch: Mixtacandona laisi 107. Borovniščica, Borovnica, HZ, 5083360 5452956, 325 m a. s. l., 21.6.02, Pascalis 2002, det. C. Meisch: Mixtacandona laisi 108. Borovniščica, Borovnica, HZ, 5084381 5452723, 320 m a. s. l., 21.6.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi 109. Borovniščica, Borovnica, HZ, 5084775 5452375, 310 m a. s. l., 8.7.02, Pascalis 2002, det. C. Meisch: Candona candida, Typhiocypris cavicola, Mktacandona chappuisi, Cypria reptans, Mktacandona laisi 110. Borovniščica, Borovnica, HZ, 5085100 5451750, 305 m a. s. l., 8.7.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandona aemonae, Mixtacandona chappuis, Mixtacandona iais, Typhiocypris cavicola, Fabaeformiscandona brevicornis 111. Borovniščica, Borovnica, HZ, 5085150 5451925, 305 m a. s. l., 8.7.02, Pascalis 2002, det. C. Meisch: Pseudocandona albicans, Typhiocypris cavicola, Mixtacandona chappuisi, Cypria reptans, Mixtacandona laisi 112. Borovniščica, Borovnica, HZ, 5086450 5451300, 300 m a. s. l., 8.7.02, Pascalis 2002, det. C. Meisch: Mixtacandona laisi, Typhlocypris cavicola, Cypria reptans, Typhlocypris cavicola 113. Borovniščica, Borovnica, HZ, 5087950 5450750, 290 m a. s. l., 8.7.02, Pascalis 2002, det. C. Meisch: Candona candida, Cypria reptans, Mktacandona chappuisi, Pseudocandona albicans 114. Podlipščica, Podlipa, HZ, 5095875 5440000, 350 m a. s. l., 9.7.02, Pascalis 2002, det. C. Meisch: Cypria lacustris, Potamocypriszschokkei 115. Podlipščica, Podlipa, HZ, 5093753 5442972, 322 m a. s. l., 16.7.02, Pascalis 2002, det. C. Meisch: Mixtacandona chappuisi, Cypria reptans 116. Podlipščica, Podlipa, HZ, 5094242 5443255, 300 m a. s. l., 17.7.02, Pascalis 2002, det. C. Meisch: Candona candida, Cypria reptans 117. Podlipščica, Podlipa, HZ, 5094169 5444172, 297 m a. s. l., 17.7.02, Pascalis 2002, det. C. Meisch: Fabaeformiscandonabreuili, Pseudocandona albicans 118. Podlipščica, Podlipa, HZ, 5094106 5445296, 293 m a. s. l., 18.7.02, Pascalis 2002, det. C. Meisch: Pseudocandona albicans 119. Podlipščica, Podlipa, HZ, 5093785 5445750, 291 m a. s. l., 18.7.02, Pascalis 2002, det. C. Meisch: Cryptocandona vavra, Pseudocandona albicans 120. Podlipščica, Podlipa, HZ, 5095243 5440925, 329 m a. s. l., 25.7.02, Pascalis 2002, det. C. Meisch: Candonopsisscourfieldi, Cryptocandona vavra, Fabaeformiscandona breuili, Typhlocypris cavicola 121. Bača, Tolmin, HZ, 5113650 5406900, 190 m a. s. l., 11.6.04, leg. N. Mori, det. C. Meisch: Pseudocandona albicans 122. Bača, Tolmin, HZ, 5113650 5406900, 190 m a. s. l., 20.11.04, leg. N. Mori, det. C. Meisch: Candona candida 123. Bača, Tolmin, HZ, 5112200 5405325, 180 m a. s. l., 11.6.04, leg. N. Mori, det. C. Meisch: Pseudocandona albicans, Candona candida 124. Bača, Tolmin, HZ, 5111916 5405126, 180 m a. s. l., 20.11.04, leg. N. Mori, det. C. Meisch: Fabaeformiscandona breuili 125. Bača, Tolmin, HZ, 5111916 5405126, 180 m a. s. l., 12.11.04, leg. N. Mori, det. C. Meisch: Cypria ophtalmica 126. Bača, Tolmin, drift, 5112200 5405325, 180 m a. s. l., 26.10.04, leg. N. Mori, det. C. Meisch: Cyclocypris ovum, Fabaeformiscandona breuili 127. Vipava, Miren, HZ, 5083614 5391029, 40 m a. s. l., 22.9.08, leg. U. Žibrat, det. N. Mori: Typhlocypris cavicola 128. Kolpa, Otok, benthos, 5052872 5524142, 140 m a. s. l., 3.8.05, leg. N. Mori, det. N. Mori: Cypridopsis vidua, Potamocypris variegata 129. Rožnik, Ljubljana, moss, 5102125 5459400, 350 m a. s. l., 15.4.05, leg. S. Lukančič, det. C. Meisch: Cyclocypris globosa, Candona neglecta, Pseudocandona lobipes 130. Cerknica Lake, Cerknica, lake littoral, 5065129 5452515, 575 m a. s. l., 1.5.05, leg. S. Lukančič, det. N. Mori: Pseudocandona pratensis 131. Bohinj Lake, Bohinjska Bistrica, lake littoral, 5126808 5410814, 527 m a. s. l., 21.6.05, leg. A. Brancelj, det. N. Mori: Bradleycyprisobliqua, Candona neglecta, Cyclocypris ovum, Cypridopsis vidua, Limnocytherina sanctipatricii 132. Bohinj Lake, Bohinjska Bistrica, lake littoral, 5127243 5414214, 527 m a. s. l., 22.6.05, leg. A. Brancelj, det. N. Mori: Candona candida, Cyclocypris ovum, Cypria ophtalmica, Pseudocandoma rostrata 133. Bohinj Lake, Bohinjska Bistrica, lake littoral, 5126884 5414222, 527 m a. s. l., 22.6.05, leg. A. Brancelj, det. N. Mori: Candona neglecta, Cyclocypris ovum, Cypria exsculpa, Cypria ophtalmica, Cypridopsis vidua, Limnocytherina sanctipatricii *HZ - hyporheic zone (depth between 30 and 60 cm in the river bed and between 60 and 90 cm in the lateral gravel bars) Discussion The intensive field sampling and taxonomical work carried out during the last ten years resulted in several new Ostracoda records for Slovenia. At present, 61 species belonging to 25 genera and 6 families are listed for Slovenia. Moreover, several new species to science need to be described. Altogether, only 23 papers have been published from 1882 until present that report on original findings of Ostracoda in Slovenian freshwaters. Since the focus of the first author's research work are groundwaters in the southern (Dinaric region) and northwestern Slovenia (Alpine region), the surface species and species that are distributed in the eastern Slovenia (Subpannonian region) are still underrepresented. The present checklist comprises species with a wide distribution across the whole of Europe as well as a number of groundwater species which are most probably endemic in Slovenia and/or the Dinaric region. In the Slovenian Red list of endangered crustaceans, five ostracod species were reported as single site endemics (C. reptans, F. aemonae, T. cavicola, T. trigonella and T. pretneri) (Sket & Brancelj 1992). After the intensive sampling of groundwater habitats (karst and alluvial aquifers) of Slovenia, it is clear that the first three species are widespread across southern and western Slovenia. The taxonomical position of the last two species is currently under revision, from which it appears that at least T. pretneri is a synonym with T. cavicola (Meisch & Mori, in preparation). Further, as proposed by Griffiths & Brancelj (1996a), the intensification of field sampling resulted in records of several species from the genera Fabaeformiscandona and Mixtacandona that had earlier not been included in a preliminary list even with a single species. It is expected that future data collection from the surface waters and from the eastern part of the country will further increase the number of recorded species and that a number of species currently considered as »single site« species will be more widespread. Povzetek Poznavanje dvoklopnikov v Sloveniji je bilo doslej razmeroma skromno. V prispevku je predstavljen najnovejši seznam recentnih vrst prostoživečih dvoklopnikov (Podocopida, Ostracoda, Crustacea) iz celinskih voda, ki vključuje podatke, povzete iz obstoječih publikacij, ter še neobjavljene najdbe za Slovenijo, pridobljene med terenskim vzorčenjem v zadnjih 10 letih. Novi seznam vsebuje 61 vrst iz 25 rodov in 6 družin. Intenzivno sistematično vzorčenje med evropskim projektom PASCALIS je pokazalo, da so nekatere vrste, predhodno znane kot lokalno endemične, splošno razširjene v slovenskem Dinarskem krasu. Pričakuje se, da je dejansko število vrst veliko večje, saj primanjkuje podatkov iz vzhodne Slovenije, ter iz površinskih voda, ki so manj intenzivno vzorčevane kot podzemne vode. Acknowledgements The field sampling was founded by the European PASCALIS project (2002-2005) and by the Slovenian Research Agency through several funding schemes. 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