©Slovenian Entomological Society, download unter www.biologiezentrum.at
LJUBLJANA, JUNIJ 2005
Vol. 13, št. 1: 9-20
NEW RECORDS OF TRUE BUGS (HETEROPTERA) OF THE BALKAN PENINSULA
Petr KMENT12), Josef BRYJA3 4) & Zdenek JINDRA5)
1} Department of Entomology, National Museum, Kunratice 1, CZ-148 00 Praha 4, Czech Republic, e-mail: sigara@post.cz 2) Department of Zoology, Faculty of Science, Charles University, Vinicna 7, CZ-128 44 Praha 2, Czech Republic 3) Department of Population Biology, Institute of Vertebrate Biology,
Academy of Sciences of the Czech Republic, CZ-675 02 Studenec 122, Czech Republic, e-mail: bryja@brno.cas.cz
4) Biodiversity Research Group, Faculty of Science, Masaryk University, Kotlarska 2, CZ-611 37 Brno, Czech Republic
5) Department of Plant Protection, Faculty of Agronomy, Czech Agricultural University, CZ-165 21 Praha 6, Czech Republic
Abstract - New records of the following species are provided: for Albania -Opisthotaenia fulvipes Reuter, 1901 and Ulmicola spinipes (Fallen, 1807), Croatia-Phytocoris scitulus scitulus Reuter, 1908, Emblethis angustus Montandon, 1890, and Proderus bellevoyei Puton, 1874, Greece - Psallus milenae Josifov, 1974, Montenegro - Arctocorisa carinata carinata (C. R. Sahlberg, 1819), Sigara limitata limitata (Fieber, 1848), Pachyxyphus lineellus (Mulsant et Rey, 1852), Berytinus striola (Ferrari, 1874), Graptostethus servus servus (Fabricius, 1787), Scolopostethus cognatus Fieber, 1861, and lschnocoris bureschi Josifov, 1976, Romania - lschnocoris bureschi, Slovenia - Scolopostethus cognatus, and European part of Turkey - Sigara limitata limitata. The occurrence of Sigara clorsalis (Leach, 1817) in Montenegro is confirmed. Additional records of rare species Brachynotocoris cyprius cyprius Wagner, 1961, Heterocapillus cavinotum Wagner, 1973, and Homoscelis ruficollis Horvath, 1884 in Greece are given.
Key words: faunistics, Heteroptera, Albania, Croatia, Greece, Macedonia, Montenegro, Romania, Slovenia, Syria, Turkey.
9
©Slovenian Entomological Society, download unter www.biologiezentrum.at
LJUBLJANA, JUNIJ 2005
Vol. 13, št. 1: 9-20
NEW RECORDS OF TRUE BUGS (HETEROPTERA) OF THE BALKAN PENINSULA
Petr KMENT12), Josef BRYJA3 4) & Zdenek JINDRA5)
1} Department of Entomology, National Museum, Kunratice 1, CZ-148 00 Praha 4, Czech Republic, e-mail: sigara@post.cz 2) Department of Zoology, Faculty of Science, Charles University, Vinicna 7, CZ-128 44 Praha 2, Czech Republic 3) Department of Population Biology, Institute of Vertebrate Biology,
Academy of Sciences of the Czech Republic, CZ-675 02 Studenec 122, Czech Republic, e-mail: bryja@brno.cas.cz
4) Biodiversity Research Group, Faculty of Science, Masaryk University, Kotlarska 2, CZ-611 37 Brno, Czech Republic
5) Department of Plant Protection, Faculty of Agronomy, Czech Agricultural University, CZ-165 21 Praha 6, Czech Republic
Abstract - New records of the following species are provided: for Albania -Opisthotaenia fulvipes Reuter, 1901 and Ulmicola spinipes (Fallen, 1807), Croatia-Phytocoris scitulus scitulus Reuter, 1908, Emblethis angustus Montandon, 1890, and Proderus bellevoyei Puton, 1874, Greece - Psallus milenae Josifov, 1974, Montenegro - Arctocorisa carinata carinata (C. R. Sahlberg, 1819), Sigara limitata limitata (Fieber, 1848), Pachyxyphus lineellus (Mulsant et Rey, 1852), Berytinus striola (Ferrari, 1874), Graptostethus servus servus (Fabricius, 1787), Scolopostethus cognatus Fieber, 1861, and lschnocoris bureschi Josifov, 1976, Romania - lschnocoris bureschi, Slovenia - Scolopostethus cognatus, and European part of Turkey - Sigara limitata limitata. The occurrence of Sigara clorsalis (Leach, 1817) in Montenegro is confirmed. Additional records of rare species Brachynotocoris cyprius cyprius Wagner, 1961, Heterocapillus cavinotum Wagner, 1973, and Homoscelis ruficollis Horvath, 1884 in Greece are given.
Key words: faunistics, Heteroptera, Albania, Croatia, Greece, Macedonia, Montenegro, Romania, Slovenia, Syria, Turkey.
9
ogiez&Gta entomologica slovenica, 13 (1), 2005
Izvleček - NOVI PODATKI O STENICAH (HETEROPTERA) BALKANSKEGA POLOTOKA
Sledeče vrste so bile prvič najdene: v Albaniji - Opisthotaenia fulvipes Reuter, 1901, in Ulmicola spinipes (Fallen, 1807), Hrvaški - Phytocoris scitulus scitulus Reuter, 1908, Emblethis angustus Montandon, 1890, in Proderus bellevoyei Puton, 1874, Grčiji -Psallus milenae Josifov, 1974, Črni gori - Arctocorisa carinata carinata (C. R. Sahlberg, 1819), Sigara limitata limitata (Fieber, 1848), Pachyxyphus lineellus (Mulsant et Rey, 1852), Beiytinus striola (Ferrari, 1874), Graptostethus servus servus (Fabricius, 1787), Scolopostethus cognatus Fieber, 1861, in Ischnocoris bureschi Josifov, 1976, Romuniji - Ischnocoris bureschi, Sloveniji - Scolopostethus cognatus, in v evropskem delu Turčije - Sigara limitata limitata. Pojavljanje vrste Sigara dorsalis (Leach, 1817) v Črni gori je potrjeno. Navedeni so novi podatki o redkih vrstah Brachynotocoris cyprius cyprius Wagner, 1961, Heterocapillus cavinotum Wagner, 1973, in Homoscelis ruficollis Horvath, 1884, v Grčiji.
Ključne besede: favnistika, Heteroptera, Albanija, Hrvaška, Grčija, Makedonija, Črna Gora, Romunija, Slovenija, Sirija, Turčija.
Introduction
True bugs of the Balkan peninsula were comprehensively treated by Josifov (1986) for the first time. Additional information on Balkan fauna was included in the monographs from the series 'Faune de France' (e.g., Moulet 1996, Pericart 1999a, 1999b, 1999c), 'Catalogue of the Heteroptera fauna of Yugoslav countries' (Protič 1998, 2001), and 'Catalogue of the Heteroptera of the Palaearctic Region' (Aukema & Rieger 1995, 1996, 1999, 2001). In spite of these valuable works, the true bug fauna of some Balkan countries still remains poorly known because of lack of local specialists and the fact that material collected by foreign collectors often remains unpublished. When determining Heteroptera from several collections deposited in the Czech Republic, we found specimens of 17 species being of particular interest. Some of them are reported here as new for the true bug faunae of Slovenia, Croatia, Montenegro, Romania, Albania, Greece, and European part of Turkey. Notes on biology are added wherever possible.
Abbreviations of collections
BMFC - coll. Beskydy Museum (Frydek-Mistek, Czech Republic); JBSC - coll. Josef Bryja (Studenec, Czech Republic); MMBC - coll. Moravian Museum (Brno, Czech Republic); PKPC - coll. Petr Kment (deposited in National Museum, Praha, Czech Republic); PMSL - coll. Slovene Museum of Natural History (Ljubljana, Slovenia); ZJPC - coll. Zdenek Jindra (Praha, Czech Republic).
List of records
Corixidae
Arctocorisa carinata carinata (C. R. Sahlberg, 1819)
10
ogiez&Gta entomologica slovenica, 13 (1), 2005
Izvleček - NOVI PODATKI O STENICAH (HETEROPTERA) BALKANSKEGA POLOTOKA
Sledeče vrste so bile prvič najdene: v Albaniji - Opisthotaenia fulvipes Reuter, 1901, in Ulmicola spinipes (Fallen, 1807), Hrvaški - Phytocoris scitulus scitulus Reuter, 1908, Emblethis angustus Montandon, 1890, in Proderus bellevoyei Puton, 1874, Grčiji -Psallus milenae Josifov, 1974, Črni gori - Arctocorisa carinata carinata (C. R. Sahlberg, 1819), Sigara limitata limitata (Fieber, 1848), Pachyxyphus lineellus (Mulsant et Rey, 1852), Beiytinus striola (Ferrari, 1874), Graptostethus servus servus (Fabricius, 1787), Scolopostethus cognatus Fieber, 1861, in Ischnocoris bureschi Josifov, 1976, Romuniji - Ischnocoris bureschi, Sloveniji - Scolopostethus cognatus, in v evropskem delu Turčije - Sigara limitata limitata. Pojavljanje vrste Sigara dorsalis (Leach, 1817) v Črni gori je potrjeno. Navedeni so novi podatki o redkih vrstah Brachynotocoris cyprius cyprius Wagner, 1961, Heterocapillus cavinotum Wagner, 1973, in Homoscelis ruficollis Horvath, 1884, v Grčiji.
Ključne besede: favnistika, Heteroptera, Albanija, Hrvaška, Grčija, Makedonija, Črna Gora, Romunija, Slovenija, Sirija, Turčija.
Introduction
True bugs of the Balkan peninsula were comprehensively treated by Josifov (1986) for the first time. Additional information on Balkan fauna was included in the monographs from the series 'Faune de France' (e.g., Moulet 1996, Pericart 1999a, 1999b, 1999c), 'Catalogue of the Heteroptera fauna of Yugoslav countries' (Protič 1998, 2001), and 'Catalogue of the Heteroptera of the Palaearctic Region' (Aukema & Rieger 1995, 1996, 1999, 2001). In spite of these valuable works, the true bug fauna of some Balkan countries still remains poorly known because of lack of local specialists and the fact that material collected by foreign collectors often remains unpublished. When determining Heteroptera from several collections deposited in the Czech Republic, we found specimens of 17 species being of particular interest. Some of them are reported here as new for the true bug faunae of Slovenia, Croatia, Montenegro, Romania, Albania, Greece, and European part of Turkey. Notes on biology are added wherever possible.
Abbreviations of collections
BMFC - coll. Beskydy Museum (Frydek-Mistek, Czech Republic); JBSC - coll. Josef Bryja (Studenec, Czech Republic); MMBC - coll. Moravian Museum (Brno, Czech Republic); PKPC - coll. Petr Kment (deposited in National Museum, Praha, Czech Republic); PMSL - coll. Slovene Museum of Natural History (Ljubljana, Slovenia); ZJPC - coll. Zdenek Jindra (Praha, Czech Republic).
List of records
Corixidae
Arctocorisa carinata carinata (C. R. Sahlberg, 1819)
10
P. Kment, J. Brya, Z. Jindra: New records of true bugs (Heteroptera) of the Balkan peninsula :entri
Montenegro or., Ivangrad env., Bjelasica Mts., small mountain lake on the ridge, 1900-2000 m, 7.VIII.2004, 1 <$, V. Syrovatka lgt., P. Kment det. (PKPC). Durmitor National Park, Jablan lake, 20.VIII.2004, 1 3 larvae, P. Bogusch & K. Svehlova lgt., P. Kment det. (PKPC).
Boreomontane species distributed in Andorra, Austria, Bulgaria, Czech Republic, Faeroe Isles, France, Great Britain (incl. Shetlands), Iceland, Italy, Liechtenstein, Norway, Russia (Central and North European Territory, Western Siberia), Slovenia, Spain, Sweden, Switzerland, and Morocco (Jansson 1995, Stys 1976, Gheit 1995, Protic 1998, Gogala 2003). According to Jansson (1979) it lives in a variety of habitats from small, more or less temporary rock pools on sea coast to permanent, relatively large and deep ponds and small lakes. In Norway it prefers large and deep lakes, but on the Atlantic coast it is also found in smaller water bodies under varying conditions, including brackish pools (Jastrey 1981). Results of experimental crosses of geographically isolated populations of A. carinata were published by Jansson (1978). New species for Montenegro.
Sigara (.Retrocorixa) limitata limitata (Fieber, 1848)
Montenegro or., Ivangrad env., Bjelasica Mts., small mountain lake on the ridge, 1900-2000 m, 7.VIII.2004, 1 1 V. Syrovatka lgt., P. Kment det. (PKPC). European Turkey, Edirne province, Edirne env. (N 41° 37' E 26° 370, flooded meadow, drain, 225 m, 16.VI.2001, 2 SS, M. Fikacek, J. Hajek & J. Straka lgt., P. Kment det (PKPC).
Eurosiberian species known from Austria, Belgium, Bosnia Hercegovina, Bulgaria, Byelorussia, Croatia, Czech Republic, Denmark, Kazakhstan (European part), Finland, France, Great Britain, Germany, Hungary, Ireland, Italy, Liechtenstein, Lithuania, Macedonia, Moldavia, Netherlands, Norway, Poland, Romania, Russia (North, Central, and South European Territory, Western Siberia), Serbia, Slovakia, ?Slovenia, Spain, Sweden, Switzerland, Ukraine, Kazakhstan (Asian part), Turkey (Asian part), and Georgia (Jansson 1995, Protic 1998, Gogala 2003). Generally rare species, preferring various types of small water habitats (e.g., Wroblewski 1980). New species for Montenegro and European part of Turkey.
Sigara (Sigara) dorsalis (Leach, 1817)
Montenegro, Vranjina env., Skadarsko jezero lake, 20.IX.2001, 1 $,3 J. Hajek lgt., P. Kment det. (PKPC).
Sigara dorsalis can unequivocally be distinguished from related species of the subgenus Sigara s. str., namely S. striata (Linnaeus, 1758) and S. basalis (A. Costa, 1843), particularly on the basis of shape of the parameres. Sigara basalis had been considered a synonym of S. dorsalis untill Rizzotti Vlach et al. (1996) restored its status of a valid species. Correspondingly, the records published from Apennine peninsula and originally referred to S. dorsalis in fact belong to S. basalis. S. dorsalis s. str. has been reported from Belgium, France, Great Britain, Greece, Ireland, Italy (northern part), Norway and Sweden (Jansson 1995, Rizzotti Vlach et al. 1996),
11
P. Kment, J. Brya, Z. Jindra: New records of true bugs (Heteroptera) of the Balkan peninsula :entri
Montenegro or., Ivangrad env., Bjelasica Mts., small mountain lake on the ridge, 1900-2000 m, 7.VIII.2004, 1 <$, V. Syrovatka lgt., P. Kment det. (PKPC). Durmitor National Park, Jablan lake, 20.VIII.2004, 1 3 larvae, P. Bogusch & K. Svehlova lgt., P. Kment det. (PKPC).
Boreomontane species distributed in Andorra, Austria, Bulgaria, Czech Republic, Faeroe Isles, France, Great Britain (incl. Shetlands), Iceland, Italy, Liechtenstein, Norway, Russia (Central and North European Territory, Western Siberia), Slovenia, Spain, Sweden, Switzerland, and Morocco (Jansson 1995, Stys 1976, Gheit 1995, Protic 1998, Gogala 2003). According to Jansson (1979) it lives in a variety of habitats from small, more or less temporary rock pools on sea coast to permanent, relatively large and deep ponds and small lakes. In Norway it prefers large and deep lakes, but on the Atlantic coast it is also found in smaller water bodies under varying conditions, including brackish pools (Jastrey 1981). Results of experimental crosses of geographically isolated populations of A. carinata were published by Jansson (1978). New species for Montenegro.
Sigara (.Retrocorixa) limitata limitata (Fieber, 1848)
Montenegro or., Ivangrad env., Bjelasica Mts., small mountain lake on the ridge, 1900-2000 m, 7.VIII.2004, 1 1 V. Syrovatka lgt., P. Kment det. (PKPC). European Turkey, Edirne province, Edirne env. (N 41° 37' E 26° 370, flooded meadow, drain, 225 m, 16.VI.2001, 2 SS, M. Fikacek, J. Hajek & J. Straka lgt., P. Kment det (PKPC).
Eurosiberian species known from Austria, Belgium, Bosnia Hercegovina, Bulgaria, Byelorussia, Croatia, Czech Republic, Denmark, Kazakhstan (European part), Finland, France, Great Britain, Germany, Hungary, Ireland, Italy, Liechtenstein, Lithuania, Macedonia, Moldavia, Netherlands, Norway, Poland, Romania, Russia (North, Central, and South European Territory, Western Siberia), Serbia, Slovakia, ?Slovenia, Spain, Sweden, Switzerland, Ukraine, Kazakhstan (Asian part), Turkey (Asian part), and Georgia (Jansson 1995, Protic 1998, Gogala 2003). Generally rare species, preferring various types of small water habitats (e.g., Wroblewski 1980). New species for Montenegro and European part of Turkey.
Sigara (Sigara) dorsalis (Leach, 1817)
Montenegro, Vranjina env., Skadarsko jezero lake, 20.IX.2001, 1 $,3 J. Hajek lgt., P. Kment det. (PKPC).
Sigara dorsalis can unequivocally be distinguished from related species of the subgenus Sigara s. str., namely S. striata (Linnaeus, 1758) and S. basalis (A. Costa, 1843), particularly on the basis of shape of the parameres. Sigara basalis had been considered a synonym of S. dorsalis untill Rizzotti Vlach et al. (1996) restored its status of a valid species. Correspondingly, the records published from Apennine peninsula and originally referred to S. dorsalis in fact belong to S. basalis. S. dorsalis s. str. has been reported from Belgium, France, Great Britain, Greece, Ireland, Italy (northern part), Norway and Sweden (Jansson 1995, Rizzotti Vlach et al. 1996),
11
©Slovenian Entomological Society, download unter www.biologiezentrum.at
Acta entomologica slovenica, 13 (1), 2005
Switzerland (Wiprachtiger 2000, Dethier 2001), and Montenegro (Grupce 1961). Josifov (1986) mentioned also Macedonia, but there is no published record from this country known to us (cf. Protic 1998). Grupce (1961) reported S. dorsalis in paper dealing with water bugs of Macedonia, he however cited only locality Skutari See (= Skadarsko jezero Lake) lying on the boundary between Montenegro and Albania. A confirmed occurrence in Montenegro.
Miridae
Brachynotocoris cyprius cyprius Wagner, 1961
Greece, W Crete, Lefka ori Mts., Floria (10 km N Kandanos), 700 m, polje, pasture and shrubs, 25.VI. 1995, 4 3 P. Lauterer lgt., J. Bryja det. (JBSC, PKPC).
The nominotypical subspecies is known from Cyprus, Iraq, Israel, Jordan (Kerzhner & Josifov 1999), and Greece: Peloponnese (Linnavuori 1999a). It lives on Olea europaea (Linnavuori 1961, 1999a). First record from Crete.
Heterocapillus cavinotum Wagner, 1973
Greece, Attiki, Poros Island, pasture, maquis, Pinus wood undergrowth, 2-50 m, 10.VI.1995, 1 S, 1 P. Lauterer lgt., J. Bryja det. (PKPC). W Crete, Lefka ori Mts., Floria (10 km N Kandanos), 700 m, polje, pasture and shrubs, 25.VI. 1995, 6 SS, 2 P. Lauterer lgt., J. Bryja det. (JBSC, PKPC). Endemic species of Greece. Originally described from Rhodes Island (Wagner 1973), and later recorded from additional localities of the Rhodes and Peloponnese peninsula (Linnavuori 1999a). The host plant is Genista acanthoclada (Linnavuori 1999a). First records from Attiki and Crete.
Opisthotaenia fulvipes Reuter, 1901
Albania, Koric env., 13.VI.1994, 1 <?, O. Hovorka lgt., J. Bryja det. (JBSC). Macedonia, Katlanovo, 14.VI.1967, 1 P. Lauterer lgt., P. Kment det. (MMBC).
Known from Bulgaria, Macedonia, Ukraine, Turkey (both European and Asian part), Armenia, Azerbaijan, and Iraq (Kerzhner & Josifov 1999). It lives on Boraginaceae: Onosma simplicissimum (Kiritshenko 1951, Putshkov 1961), Echium, and Lithospermum (Seidenstucker 1968). New species for Albania and additional record for Macedonia (for published Macedonian localities see Protic (1998)).
Pachyxyphus lineellus (Mulsant et Rey, 1852)
Montenegro occ., Budva (6 km NW), 5.VI.1967, 1 P. Lauterer lgt., P. Kment det. (MMBC).
Mediterranean species occurring in Albania, Bosnia Hercegovina, Bulgaria, Croatia, France, Greece, Italy, Portugal, Spain, Ukraine; Turkey (both European and Asian parts), Cyprus, Georgia, Israel, Jordan; Algeria, Morocco, and Tunisia
68
©Slovenian Entomological Society, download unter www.biologiezentrum.at
Acta entomologica slovenica, 13 (1), 2005
Switzerland (Wiprachtiger 2000, Dethier 2001), and Montenegro (Grupce 1961). Josifov (1986) mentioned also Macedonia, but there is no published record from this country known to us (cf. Protic 1998). Grupce (1961) reported S. dorsalis in paper dealing with water bugs of Macedonia, he however cited only locality Skutari See (= Skadarsko jezero Lake) lying on the boundary between Montenegro and Albania. A confirmed occurrence in Montenegro.
Miridae
Brachynotocoris cyprius cyprius Wagner, 1961
Greece, W Crete, Lefka ori Mts., Floria (10 km N Kandanos), 700 m, polje, pasture and shrubs, 25.VI. 1995, 4 3 P. Lauterer lgt., J. Bryja det. (JBSC, PKPC).
The nominotypical subspecies is known from Cyprus, Iraq, Israel, Jordan (Kerzhner & Josifov 1999), and Greece: Peloponnese (Linnavuori 1999a). It lives on Olea europaea (Linnavuori 1961, 1999a). First record from Crete.
Heterocapillus cavinotum Wagner, 1973
Greece, Attiki, Poros Island, pasture, maquis, Pinus wood undergrowth, 2-50 m, 10.VI.1995, 1 S, 1 P. Lauterer lgt., J. Bryja det. (PKPC). W Crete, Lefka ori Mts., Floria (10 km N Kandanos), 700 m, polje, pasture and shrubs, 25.VI. 1995, 6 SS, 2 P. Lauterer lgt., J. Bryja det. (JBSC, PKPC). Endemic species of Greece. Originally described from Rhodes Island (Wagner 1973), and later recorded from additional localities of the Rhodes and Peloponnese peninsula (Linnavuori 1999a). The host plant is Genista acanthoclada (Linnavuori 1999a). First records from Attiki and Crete.
Opisthotaenia fulvipes Reuter, 1901
Albania, Koric env., 13.VI.1994, 1 <?, O. Hovorka lgt., J. Bryja det. (JBSC). Macedonia, Katlanovo, 14.VI.1967, 1 P. Lauterer lgt., P. Kment det. (MMBC).
Known from Bulgaria, Macedonia, Ukraine, Turkey (both European and Asian part), Armenia, Azerbaijan, and Iraq (Kerzhner & Josifov 1999). It lives on Boraginaceae: Onosma simplicissimum (Kiritshenko 1951, Putshkov 1961), Echium, and Lithospermum (Seidenstucker 1968). New species for Albania and additional record for Macedonia (for published Macedonian localities see Protic (1998)).
Pachyxyphus lineellus (Mulsant et Rey, 1852)
Montenegro occ., Budva (6 km NW), 5.VI.1967, 1 P. Lauterer lgt., P. Kment det. (MMBC).
Mediterranean species occurring in Albania, Bosnia Hercegovina, Bulgaria, Croatia, France, Greece, Italy, Portugal, Spain, Ukraine; Turkey (both European and Asian parts), Cyprus, Georgia, Israel, Jordan; Algeria, Morocco, and Tunisia
68
P. Kment, J. Brya, Z. Jinclra: New records of true bugs (Heteroptera) of the Balkan peninsula :en
(Kerzhner & Josifov 1999, Carapezza 2002). It lives on Cistus species (Wagner 1975). New species for Montenegro.
Phytocoris (Exophytocoris) scitulus scitulus Reuter, 1908
Croatia, Petrcane, 6 km NW of Zadar, on Tamarix, 16.VII.2000, 1 M. Mantic lgt., J. Bryja det. (JBSC).
The nominotypical subspecies is distributed in Bulgaria, Moldavia, Russia (Caucasus), Georgia, Armenia, Azerbaijan (Kerzhner & Josifov 1999), and Iran (Linnavuori 1999b, Linnavuori & Modarres 1999). The subspecies P s. 'frater Kerzhner, 1964 is endemic to Crimea (Ukraine). According to Putshkov & Putshkov (1983), P scitulus (subspecies not distinguished) lives on many leafy shrubs and trees, often on Crataegus sp. and Mains sp., especially if they are infested with psyllids. Linnavuori (1962) reported it from Taxus baccata, Linnavuori (1999b) from Cupressus, Juniperus polycarpus and Olea europaea. New species for Croatia.
Psallus (Psallus) milenae Josifov, 1974
Greece, NE Peloponissos, Korinthia distr., Drevenakia (9 km N of Mykenai), 200 m, pasture & maquis with Pyrus amygclaliformis, Pistacia lentiscus, and Quercus coccifera, 6.V.1999, 3 SS, 1 P. Lauterer lgt., J. Bryja det. (JBSC).
Known from Bulgaria, Macedonia, Asian part of Turkey (Josifov 1974, 1986, Wagner & Weber 1978; Kerzhner & Josifov 1999). According to Josifov (1974), it lives on Quercus pubescens. Our specimens were most probably collected from Quercus coccifera. New species for Greece.
Berytidae
Berytinus (Lizinus) striola (Ferrari, 1874)
Montenegro occ., Djurasevici —► Delfin, maquis, cemetery, 50-150 m, 25.X. 1982, 2 P. Lauterer lgt., J. L. Stehlik det. (MMBC).
Mediterranean species occurring in Albania, Austria, Bosnia Hercegovina, Bulgaria, Croatia, Czech Republic, France, Greece, Hungary, Italy, Portugal, Romania, Russia (South European Territory), Serbia, Slovakia, Slovenia, Spain, Ukraine; Algeria, Morocco, Tunisia; Armenia, Turkey (Asian part), Cyprus, Georgia, Iraq, Israel, Lebanon, and Syria (Pericart 2001a, Protic 2001). Josifov (1986) and Protic (1999) listed also Macedonia. Biology of this species is poorly known (see Pericart 1984). In Moravia (Czech Republic) it was collected on xerothermic sites on various substrates (limestone, granodiorites, diorites, loamy substrate, rarely also on sand) where it feeds on Fabaceae (Stehlik & Vavrinova 1990). New species for Montenegro.
Lygaeidae
Graptostethus servus servus (Fabricius, 1787)
13
P. Kment, J. Brya, Z. Jinclra: New records of true bugs (Heteroptera) of the Balkan peninsula :en
(Kerzhner & Josifov 1999, Carapezza 2002). It lives on Cistus species (Wagner 1975). New species for Montenegro.
Phytocoris (Exophytocoris) scitulus scitulus Reuter, 1908
Croatia, Petrcane, 6 km NW of Zadar, on Tamarix, 16.VII.2000, 1 M. Mantic lgt., J. Bryja det. (JBSC).
The nominotypical subspecies is distributed in Bulgaria, Moldavia, Russia (Caucasus), Georgia, Armenia, Azerbaijan (Kerzhner & Josifov 1999), and Iran (Linnavuori 1999b, Linnavuori & Modarres 1999). The subspecies P s. 'frater Kerzhner, 1964 is endemic to Crimea (Ukraine). According to Putshkov & Putshkov (1983), P scitulus (subspecies not distinguished) lives on many leafy shrubs and trees, often on Crataegus sp. and Mains sp., especially if they are infested with psyllids. Linnavuori (1962) reported it from Taxus baccata, Linnavuori (1999b) from Cupressus, Juniperus polycarpus and Olea europaea. New species for Croatia.
Psallus (Psallus) milenae Josifov, 1974
Greece, NE Peloponissos, Korinthia distr., Drevenakia (9 km N of Mykenai), 200 m, pasture & maquis with Pyrus amygclaliformis, Pistacia lentiscus, and Quercus coccifera, 6.V.1999, 3 SS, 1 P. Lauterer lgt., J. Bryja det. (JBSC).
Known from Bulgaria, Macedonia, Asian part of Turkey (Josifov 1974, 1986, Wagner & Weber 1978; Kerzhner & Josifov 1999). According to Josifov (1974), it lives on Quercus pubescens. Our specimens were most probably collected from Quercus coccifera. New species for Greece.
Berytidae
Berytinus (Lizinus) striola (Ferrari, 1874)
Montenegro occ., Djurasevici —► Delfin, maquis, cemetery, 50-150 m, 25.X. 1982, 2 P. Lauterer lgt., J. L. Stehlik det. (MMBC).
Mediterranean species occurring in Albania, Austria, Bosnia Hercegovina, Bulgaria, Croatia, Czech Republic, France, Greece, Hungary, Italy, Portugal, Romania, Russia (South European Territory), Serbia, Slovakia, Slovenia, Spain, Ukraine; Algeria, Morocco, Tunisia; Armenia, Turkey (Asian part), Cyprus, Georgia, Iraq, Israel, Lebanon, and Syria (Pericart 2001a, Protic 2001). Josifov (1986) and Protic (1999) listed also Macedonia. Biology of this species is poorly known (see Pericart 1984). In Moravia (Czech Republic) it was collected on xerothermic sites on various substrates (limestone, granodiorites, diorites, loamy substrate, rarely also on sand) where it feeds on Fabaceae (Stehlik & Vavrinova 1990). New species for Montenegro.
Lygaeidae
Graptostethus servus servus (Fabricius, 1787)
13
©Slovenian Entomological Society, download unter www.biologiezentrum.at
Acta entomologica slovenica, 13 (1), 2005
Montenegro occ., Radovici, Plavi Horizont, maquis, 3-20 m, 13.-15.X.1982, 1 P. Lauterer lgt., P. Kment det. (MMBC).
A species widely distributed in the Old World, reaching its northernmost limit in Mediterranean: recorded from Albania, Croatia, France, Greece, Italy, Portugal, Serbia, Spain; Algeria, Egypt, Morocco; Turkey (Asian part), China (Southeastern and Southwestern Territories, Western Plateau), Cyprus, Israel, Japan, Korea, Saudi Arabia, Syria, Tadjikistan, Yemen; Afrotropical and Oriental Regions, Australia, Polynesia, and Hawaii (Pericart 2001b, Protic 2001). A polyphagous species that lives on ground or the basis of vegetation and prefers field cultures and orchards. In the tropical parts of its distributional range it is known as a pest of several crops (e.g., Gossypium, Sorghum vulgare, Ipomoea tuberosa) (see Pericart (1999a) for details). New species for Montenegro.
Rhyparochromidae
Emblethis angustus Montandon, 1890
Croatia occ., Brae Island mer., Bol env., environs of Gazul Suma under Vidova Gora Mt. (43°18'N 16°38'E), 700-740 m, pastures, on ground, 7.IX.2002, 1 S, P. Kment lgt. et det. (PKPC); dtto, Vidova Gora Mt. (43°16'N 16°39'E), 740 m, pastures under the top, under stones, 10.IX.2002, 1 <S, 2 P. Kment lgt. et det. (PKPC).
Mediterranean and Turanian species known from Bulgaria, France, Greece, Italy, Malta, Macedonia, Portugal, Russia (South European Territory), Spain, Ukraine; Algeria, Canary Islands, Egypt, Lybia, Madeira, Morocco, Tunisia; Azerbaijan, Afghanistan, Kazakhstan (Asian part), Turkey (Asian part), Georgia, Iran, Iraq, Israel, Syria, Tadjikistan, Turkmenistan, and Uzbekistan. Outside Palaearctic region it occurs also on Cape Verde Islands (Pericart 2001b). Pericart (1999c) did not give any infornation on biology of E. angustus. Our Croatian specimens were collected on shortly grazed pastures on limestone, usually under stones. New species for Croatia.
Ischnocoris bureschi Josifov, 1976
Montenegro, Becici near Budva, 4.V. 1975, 1 2 J. Strejcek lgt., Z. Jindra det. (ZJPC). Romania mer., Bucura, 22.V. 1993, 1 S (brachypterous), O. Majzlan lgt., P. Kment det. (BMFC).
Described from Bulgaria (Josifov 1976), and subsequently recorded from Albania, Greece, Croatia, Macedonia and Asian part of Turkey (Pericart 1999b, 2001b). Biology of this species is unknown. New species for Montenegro and Romania.
Homoscelis ruficollis Horvath, 1884
Greece occ., Kerkira (= Corfu) Island bor., Kassiopi (39°47'44"N 19°55' 09" E), 3.V.2002, 1 1 F. St'ahlavsky lgt., P. Kment det. (PKPC).
68
©Slovenian Entomological Society, download unter www.biologiezentrum.at
Acta entomologica slovenica, 13 (1), 2005
Montenegro occ., Radovici, Plavi Horizont, maquis, 3-20 m, 13.-15.X.1982, 1 P. Lauterer lgt., P. Kment det. (MMBC).
A species widely distributed in the Old World, reaching its northernmost limit in Mediterranean: recorded from Albania, Croatia, France, Greece, Italy, Portugal, Serbia, Spain; Algeria, Egypt, Morocco; Turkey (Asian part), China (Southeastern and Southwestern Territories, Western Plateau), Cyprus, Israel, Japan, Korea, Saudi Arabia, Syria, Tadjikistan, Yemen; Afrotropical and Oriental Regions, Australia, Polynesia, and Hawaii (Pericart 2001b, Protic 2001). A polyphagous species that lives on ground or the basis of vegetation and prefers field cultures and orchards. In the tropical parts of its distributional range it is known as a pest of several crops (e.g., Gossypium, Sorghum vulgare, Ipomoea tuberosa) (see Pericart (1999a) for details). New species for Montenegro.
Rhyparochromidae
Emblethis angustus Montandon, 1890
Croatia occ., Brae Island mer., Bol env., environs of Gazul Suma under Vidova Gora Mt. (43°18'N 16°38'E), 700-740 m, pastures, on ground, 7.IX.2002, 1 S, P. Kment lgt. et det. (PKPC); dtto, Vidova Gora Mt. (43°16'N 16°39'E), 740 m, pastures under the top, under stones, 10.IX.2002, 1 <S, 2 P. Kment lgt. et det. (PKPC).
Mediterranean and Turanian species known from Bulgaria, France, Greece, Italy, Malta, Macedonia, Portugal, Russia (South European Territory), Spain, Ukraine; Algeria, Canary Islands, Egypt, Lybia, Madeira, Morocco, Tunisia; Azerbaijan, Afghanistan, Kazakhstan (Asian part), Turkey (Asian part), Georgia, Iran, Iraq, Israel, Syria, Tadjikistan, Turkmenistan, and Uzbekistan. Outside Palaearctic region it occurs also on Cape Verde Islands (Pericart 2001b). Pericart (1999c) did not give any infornation on biology of E. angustus. Our Croatian specimens were collected on shortly grazed pastures on limestone, usually under stones. New species for Croatia.
Ischnocoris bureschi Josifov, 1976
Montenegro, Becici near Budva, 4.V. 1975, 1 2 J. Strejcek lgt., Z. Jindra det. (ZJPC). Romania mer., Bucura, 22.V. 1993, 1 S (brachypterous), O. Majzlan lgt., P. Kment det. (BMFC).
Described from Bulgaria (Josifov 1976), and subsequently recorded from Albania, Greece, Croatia, Macedonia and Asian part of Turkey (Pericart 1999b, 2001b). Biology of this species is unknown. New species for Montenegro and Romania.
Homoscelis ruficollis Horvath, 1884
Greece occ., Kerkira (= Corfu) Island bor., Kassiopi (39°47'44"N 19°55' 09" E), 3.V.2002, 1 1 F. St'ahlavsky lgt., P. Kment det. (PKPC).
68
P. Kment, J. Brya, Z. Jindra: New records of true bugs (Heteroptera) of the Balkan peninsula :entri
H. ruficollis was described from Kerkira (= Corfu) Island by Horvath (1884). Linnavuori (1953) recorded it from Corfu again (two specimens collected in 1896), and also from the Lesbos Island (one specimen collected in 1904). Few other localities are known from the Asian part of Turkey (Hoberlandt 1956, Pericart 1999b). Biology is poorly known (see Pericart 1999b). Seidenstiicker (1960) gave the following characteristics of the Turkish localities: 'On wet soil. Near Bursa among group of Castanea sp. under ferns on the red loamy ground of a spring horizont. Near Golba§i in floodplain forest (Populus) under Genista sp. at a lake margin' Our specimens are the first ones collected in Greece after nearly a century.
Proderus bellevoyei Puton, 1874
Croatia occ., Brae Island or., Sumartin (43°17'N 16°52'E), 40 m, abandoned gardens at the village, under stone, 5.IX.2002, 1 P. Kment & M. Horsak lgt., P. Kment det. (PKPC). Additional records: Greece occ., Kerkira (= Corfu) Island bor., Agrafoi (39°45'N 19°44'E), 8.V.2002, 1 F. St'ahlavsky lgt., P. Kment det. (PKPC). Syria bor. occ., Horns env., castle Qalaat al Hosn, 30.III.-4.IV.2001, 1 M. Rezac lgt., P. Kment det. (PKPC). Turkey mer., Antalya province, Beldibi near Kemer, 1.-17.VII.1998, 1 -, P. Bulirsch lgt., Z. Jindra det. (ZJPC). Turkey occ., Izmir province, Antique Ephesos, 29.IV.1991, 1 S, 1 Z. Jindra lgt. et det. (ZJPC). Turkey or., Sivas province (39°53'N 37°37'E), Demiryurt, Todiirge Golii lake env., gypsum hills, rocky steppe, in litter, 28.VI.2002. 1 P. Kment lgt. et det. (PKPC).
East-Mediterranean species known from scattered localities in Albania, Bosnia Herzegovina, Bulgaria, Greece, Macedonia, Russia (Daghestan), Ukraine (Crimea); Azerbaijan (incl. Nakhichevan), Armenia, Asian part of Turkey, Cyprus, Israel, Jordan, Lebanon, and Syria (Pericart 1999b, 2001b). It was described from Syria, but there is no published exact Syrian locality (Pericart 1999b). The biology is almost unknown (cf. Pericart 1999c). Kiritshenko (1908, as P crassicornis Jakovlev) reported its finding under stones along a moist garden wall in Crimea (Ukraine), Putshkov (1969) mentioned other Crimean localities, including those on the sea coast, but without ecological details. In Nakhichevan it was recorded under stones on dry slopes (Putshkov 1969). The condition of our Croatian and Turkish (Sivas) records are similar to those characteristics. Adults occur from March to late autumn (Putschkov 1969, our records), not from September to late autumn as erroneously indicated by Pericart (1999c). New species for Croatia and first exactly located record from Syria after the description.
Scolopostethus cognatus Fieber, 1861
Montenegro, Lovcen, Stirovnik, 1 km SW, polje and hill, 1300-1400 m, 20.X. 1982, 2 P. Lauterer lgt., J. L. Stehlik det., P. Kment revid. (MMBC). Slovenia mer. occ., Kozina env., Kastelec, 30.IV.2002, Z. Malinka lgt., P. Kment det. (PMSL).
Mediterranean species distributed in Albania, Bulgaria, Croatia, France, Greece, Hungary, Italy, Macedonia, Portugal, Romania, Spain, Switzerland, Tunisia, and
11
P. Kment, J. Brya, Z. Jindra: New records of true bugs (Heteroptera) of the Balkan peninsula :entri
H. ruficollis was described from Kerkira (= Corfu) Island by Horvath (1884). Linnavuori (1953) recorded it from Corfu again (two specimens collected in 1896), and also from the Lesbos Island (one specimen collected in 1904). Few other localities are known from the Asian part of Turkey (Hoberlandt 1956, Pericart 1999b). Biology is poorly known (see Pericart 1999b). Seidenstiicker (1960) gave the following characteristics of the Turkish localities: 'On wet soil. Near Bursa among group of Castanea sp. under ferns on the red loamy ground of a spring horizont. Near Golba§i in floodplain forest (Populus) under Genista sp. at a lake margin' Our specimens are the first ones collected in Greece after nearly a century.
Proderus bellevoyei Puton, 1874
Croatia occ., Brae Island or., Sumartin (43°17'N 16°52'E), 40 m, abandoned gardens at the village, under stone, 5.IX.2002, 1 P. Kment & M. Horsak lgt., P. Kment det. (PKPC). Additional records: Greece occ., Kerkira (= Corfu) Island bor., Agrafoi (39°45'N 19°44'E), 8.V.2002, 1 F. St'ahlavsky lgt., P. Kment det. (PKPC). Syria bor. occ., Horns env., castle Qalaat al Hosn, 30.III.-4.IV.2001, 1 M. Rezac lgt., P. Kment det. (PKPC). Turkey mer., Antalya province, Beldibi near Kemer, 1.-17.VII.1998, 1 -, P. Bulirsch lgt., Z. Jindra det. (ZJPC). Turkey occ., Izmir province, Antique Ephesos, 29.IV.1991, 1 S, 1 Z. Jindra lgt. et det. (ZJPC). Turkey or., Sivas province (39°53'N 37°37'E), Demiryurt, Todiirge Golii lake env., gypsum hills, rocky steppe, in litter, 28.VI.2002. 1 P. Kment lgt. et det. (PKPC).
East-Mediterranean species known from scattered localities in Albania, Bosnia Herzegovina, Bulgaria, Greece, Macedonia, Russia (Daghestan), Ukraine (Crimea); Azerbaijan (incl. Nakhichevan), Armenia, Asian part of Turkey, Cyprus, Israel, Jordan, Lebanon, and Syria (Pericart 1999b, 2001b). It was described from Syria, but there is no published exact Syrian locality (Pericart 1999b). The biology is almost unknown (cf. Pericart 1999c). Kiritshenko (1908, as P crassicornis Jakovlev) reported its finding under stones along a moist garden wall in Crimea (Ukraine), Putshkov (1969) mentioned other Crimean localities, including those on the sea coast, but without ecological details. In Nakhichevan it was recorded under stones on dry slopes (Putshkov 1969). The condition of our Croatian and Turkish (Sivas) records are similar to those characteristics. Adults occur from March to late autumn (Putschkov 1969, our records), not from September to late autumn as erroneously indicated by Pericart (1999c). New species for Croatia and first exactly located record from Syria after the description.
Scolopostethus cognatus Fieber, 1861
Montenegro, Lovcen, Stirovnik, 1 km SW, polje and hill, 1300-1400 m, 20.X. 1982, 2 P. Lauterer lgt., J. L. Stehlik det., P. Kment revid. (MMBC). Slovenia mer. occ., Kozina env., Kastelec, 30.IV.2002, Z. Malinka lgt., P. Kment det. (PMSL).
Mediterranean species distributed in Albania, Bulgaria, Croatia, France, Greece, Hungary, Italy, Macedonia, Portugal, Romania, Spain, Switzerland, Tunisia, and
11
©Slovenian Entomological Society, download unter www.biologiezentrum.at
Acta entomologica slovenica, 13 (1), 2005
?Turkey (Asian part) (Pericart 2001b). Pericart (1999b) did not include any information on its biology. The Hoberlandt's (1956) specimen was found on the ground in wooded land formation. Josifov (1964) characterized its habitat as moist detritus on river banks. According to Rampazzi & Dethier (1997) it lives on heathlands of Calluna vulgaris; they found one specimen on margin of peatbog and mixed broadleaved forest with predominant Castanea sativa. New species for Slovenia and Montenegro.
Coreidae
Ulmicola spinipes (Fallen, 1807)
Albania bor., Boge env. (42° 24'N 19° 38' E), 19.-21.VI.1994, 1 O. Hovorka lgt., P. Kment det. (PKPC).
A species of European distribution, known from Austria, Bulgaria, Czech Republic, Finland, France, Germany, Hungary, Italy, Macedonia, Norway, Poland, Romania, Slovakia, Slovenia, Spain, Sweden, and Ukraine (Moulet 1995). Josifov (1986) considered it as a boreomontane Eurosiberian element. U. spinipes is known from various host plants: Trifolium, Melilotus, Medicago, Lamium, Chenopodium, Alnus, Carpinus (Moulet 1995). In southern Moravia (Czech Republic) an adult was found on a dry cadaver (head of Ovis musimon) together with Ceraleptus lividus Stein, 1858 (Coreidae). New species for Albania.
Acknowledgements
We thank Jiff Hajek (National Museum, Praha), Pavel Lauterer and Igor Malenovsky (both Moravina Museum, Brno), Jitka Vilimova, Petr Bogusch, and Martin Fikacek (all Charles University, Praha), Magdalena Rohacova (Beskydy Museum, Frydek-Mistek), Marion Mantic (Hlucin), Zdenek Malinka (Opava), and Vit Syrovatka (Masaryk University, Brno) for the possibility to study their collections or the gifts of their material. We are also indebted to Igor Malenovsky for help with papers in French and critical reading of the manuscript, and to Michal Horsak (Masaryk University, Brno) for assistance in the field. Our work was partly supported by grants from the Ministry of Education MSMT 0021620828 (to Charles University, Praha) and MSMT 0021622416 (to Masaryk University, Brno), and grant from the Ministry of Culture MK 00002327201 (to National Museum, Praha).
References
Aukema, B. & Rieger, Ch. (eds.), 1995: Catalogue of the Heteroptera of the Palaearctic Region. Vol. 1. Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha. xxvi + 222 pp. The Netherlands Entomological Society, Amsterdam.
68
©Slovenian Entomological Society, download unter www.biologiezentrum.at
Acta entomologica slovenica, 13 (1), 2005
?Turkey (Asian part) (Pericart 2001b). Pericart (1999b) did not include any information on its biology. The Hoberlandt's (1956) specimen was found on the ground in wooded land formation. Josifov (1964) characterized its habitat as moist detritus on river banks. According to Rampazzi & Dethier (1997) it lives on heathlands of Calluna vulgaris; they found one specimen on margin of peatbog and mixed broadleaved forest with predominant Castanea sativa. New species for Slovenia and Montenegro.
Coreidae
Ulmicola spinipes (Fallen, 1807)
Albania bor., Boge env. (42° 24'N 19° 38' E), 19.-21.VI.1994, 1 O. Hovorka lgt., P. Kment det. (PKPC).
A species of European distribution, known from Austria, Bulgaria, Czech Republic, Finland, France, Germany, Hungary, Italy, Macedonia, Norway, Poland, Romania, Slovakia, Slovenia, Spain, Sweden, and Ukraine (Moulet 1995). Josifov (1986) considered it as a boreomontane Eurosiberian element. U. spinipes is known from various host plants: Trifolium, Melilotus, Medicago, Lamium, Chenopodium, Alnus, Carpinus (Moulet 1995). In southern Moravia (Czech Republic) an adult was found on a dry cadaver (head of Ovis musimon) together with Ceraleptus lividus Stein, 1858 (Coreidae). New species for Albania.
Acknowledgements
We thank Jiff Hajek (National Museum, Praha), Pavel Lauterer and Igor Malenovsky (both Moravina Museum, Brno), Jitka Vilimova, Petr Bogusch, and Martin Fikacek (all Charles University, Praha), Magdalena Rohacova (Beskydy Museum, Frydek-Mistek), Marion Mantic (Mucin), Zdenek Malinka (Opava), and Vit Syrovatka (Masaryk University, Brno) for the possibility to study their collections or the gifts of their material. We are also indebted to Igor Malenovsky for help with papers in French and critical reading of the manuscript, and to Michal Horsak (Masaryk University, Brno) for assistance in the field. Our work was partly supported by grants from the Ministry of Education MSMT 0021620828 (to Charles University, Praha) and MSMT 0021622416 (to Masaryk University, Brno), and grant from the Ministry of Culture MK 00002327201 (to National Museum, Praha).
References
Aukema, B. & Rieger, Ch. (eds.), 1995: Catalogue of the Heteroptera of the Palaearctic Region. Vol. 1. Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha. xxvi + 222 pp. The Netherlands Entomological Society, Amsterdam.
68
©Slovenian Entomological Society, download unter www.biologiezentrum.at
P. Kment, J. Brya, Z. Jindra: New records of true bugs (Heteroptera) of the Balkan peninsula
Aukema, B. & Rieger, Ch. (eds.) 1996: Catalogue of the Heteroptera of the Palaearctic Region. Vol. 2. Cimicomorpha I. xiv + 361 pp. The Netherlands Entomological Society, Amsterdam.
Aukema, B. & Rieger, Ch. (eds.) 1999: Catalogue of the Heteroptera of the Palaearctic Region. Vol. 3. Cimicomorpha II. xiv + 577 pp. The Netherlands Entomological Society, Amsterdam.
Aukema, B. & Rieger, Ch. (eds.) 2001: Catalogue of the Heteroptera of the Palaearctic Region. Vol. 4. Pentatomomorpha I. xiv + 346 pp. The Netherlands Entomological Society, Amsterdam.
Carapezza, A., 2002: Heteroptera of Jordan: new taxa and new records. Naturalista Siciliano, 26: 35-76.
Dethier, M., 2001: Présence de Sigara dorsalis (Leach, 1817) en Suisse (Heteroptera Corixidae). Bulletin Romand d'Entomologie, 19: 17-21.
Gheit, A., 1995: Catalogue des Hémiptères Hydrocorises et Amphibiocorises des provinces nord Marocaines (lre note). Entomologiste (Paris), 51: 241-249.
Gogala, A., 2003: Heteroptera of Slovenia, I.: Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha. Annals for Istrian and Mediterranean Studies, Series Historia Naturalis, 13: 229-240.
Grupce, R., 1961: Beitrag zur Kenntnis der Heteroptera aquatica (Corixinae) von Mazedonien. Fragmenta Balcanica, 4: 29-36.
Hoberlandt, L., 1956: Results of the Zoological Scientific Expedition of the National Museum in Prague to Turkey. 18. Hemiptera IV. Terrestrial Hemiptera-Heteroptera of Turkey. Acta Entomologica Musei Nationalis Pragae, Suppl., 3(1955): 1-264.
Jansson, A., 1978: Viability of progeny in experimental crosses between geographically isolated populations of Arctocorisa carinata (C. Sahlberg) (Heteroptera, Corixidae). Annales Zoologici Fennici, 15: 77-83.
Jansson, A., 1979: Reproductive isolation and experimental hybridization between Arctocorisa carinata and A. germari (Heteroptera, Corixidae). Annales Zoologici Fennici, 16: 89-104.
Jansson, A., 1995: Family Corixidae Leach, 1815 - water boatmen, pp. 26-56. In: Aukema B. & Rieger Ch. (eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 1. Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha. xxvi + 222 pp. The Netherlands Entomological Society, Amsterdam.
Jastrey, J. T., 1981: Distribution and ecology of Norwegian water-bugs (Hem., Heteroptera). Fauna Norvegica, Series B, 28: 1-24.
Josifov, M., 1964: Vidov s'stav i razprostranenie na nasekomite ot razred Heteroptera v B'lgariya, chast II. (Artbestand und Verbreitung der Insekten von der Ordnung Heteroptera in Bulgarien, Teil II.). Izvestiya na Zoologicheskiya Institut s Muzey (Sofia), 16: 83-150.
Josifov, M., 1974: Eine neue Psallus-Art aus Bulgarien und eine neue Orthotylus-Art aus Kirgisien (Heteroptera, Miridae). Reichenbachia, 15: 89-92.
17
©Slovenian Entomological Society, download unter www.biologiezentrum.at
P. Kment, J. Brya, Z. Jindra: New records of true bugs (Heteroptera) of the Balkan peninsula
Aukema, B. & Rieger, Ch. (eds.) 1996: Catalogue of the Heteroptera of the Palaearctic Region. Vol. 2. Cimicomorpha I. xiv + 361 pp. The Netherlands Entomological Society, Amsterdam.
Aukema, B. & Rieger, Ch. (eds.) 1999: Catalogue of the Heteroptera of the Palaearctic Region. Vol. 3. Cimicomorpha II. xiv + 577 pp. The Netherlands Entomological Society, Amsterdam.
Aukema, B. & Rieger, Ch. (eds.) 2001: Catalogue of the Heteroptera of the Palaearctic Region. Vol. 4. Pentatomomorpha I. xiv + 346 pp. The Netherlands Entomological Society, Amsterdam.
Carapezza, A., 2002: Heteroptera of Jordan: new taxa and new records. Naturalista Siciliano, 26: 35-76.
Dethier, M., 2001: Présence de Sigara dorsalis (Leach, 1817) en Suisse (Heteroptera Corixidae). Bulletin Romand d'Entomologie, 19: 17-21.
Gheit, A., 1995: Catalogue des Hémiptères Hydrocorises et Amphibiocorises des provinces nord Marocaines (lre note). Entomologiste (Paris), 51: 241-249.
Gogala, A., 2003: Heteroptera of Slovenia, I.: Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha. Annals for Istrian and Mediterranean Studies, Series Historia Naturalis, 13: 229-240.
Grupce, R., 1961: Beitrag zur Kenntnis der Heteroptera aquatica (Corixinae) von Mazedonien. Fragmenta Balcanica, 4: 29-36.
Hoberlandt, L., 1956: Results of the Zoological Scientific Expedition of the National Museum in Prague to Turkey. 18. Hemiptera IV. Terrestrial Hemiptera-Heteroptera of Turkey. Acta Entomologica Musei Nationalis Pragae, Suppl., 3(1955): 1-264.
Jansson, A., 1978: Viability of progeny in experimental crosses between geographically isolated populations of Arctocorisa carinata (C. Sahlberg) (Heteroptera, Corixidae). Annales Zoologici Fennici, 15: 77-83.
Jansson, A., 1979: Reproductive isolation and experimental hybridization between Arctocorisa carinata and A. germari (Heteroptera, Corixidae). Annales Zoologici Fennici, 16: 89-104.
Jansson, A., 1995: Family Corixidae Leach, 1815 - water boatmen, pp. 26-56. In: Aukema B. & Rieger Ch. (eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 1. Enicocephalomorpha, Dipsocoromorpha, Nepomorpha, Gerromorpha and Leptopodomorpha. xxvi + 222 pp. The Netherlands Entomological Society, Amsterdam.
Jastrey, J. T., 1981: Distribution and ecology of Norwegian water-bugs (Hem., Heteroptera). Fauna Norvegica, Series B, 28: 1-24.
Josifov, M., 1964: Vidov s'stav i razprostranenie na nasekomite ot razred Heteroptera v B'lgariya, chast II. (Artbestand und Verbreitung der Insekten von der Ordnung Heteroptera in Bulgarien, Teil II.). Izvestiya na Zoologicheskiya Institut s Muzey (Sofia), 16: 83-150.
Josifov, M., 1974: Eine neue Psallus-Art aus Bulgarien und eine neue Orthotylus-Art aus Kirgisien (Heteroptera, Miridae). Reichenbachia, 15: 89-92.
17
©Slovenian Entomological Society, download unter www.biologiezentrum.at
Acta entomologica slovenica, 13 (1), 2005
Josifov, M., 1976: Eine neue Lygaeidae-Art (Heteroptera) aus der Balkanhalbinsel. Acta Zoologica Bulgarica, 4: 65-66.
Josifov, M., 1986: Verzeichnis der von der Balkanhalbinsel bekannten Heteropterenarten (Insecta, Heteroptera). Faunistische Abhandlungen Staatliches Museum für Tierkunde Dresden, 14: 61-93.
Kerzhner, I. M. & Josifov, M., 1999: Miridae Hahn 1833, pp. 1-577. In: Aukema B. & Rieger Ch. (eds.): Catalogue of the Heteroptera of the Palaearctic Region. Vol. 3. Cimicomorpha II. xiv + 577 pp. The Netherlands Entomological Society, Amsterdam.
Kiritshenko, A. N., 1908: K faune Hemiptera - Heteroptera Kryma. [To the Hemiptera - Heteroptera fauna of Crimea]. Russkoe Entomologicheskoe Obozrenie, 8: 234-239 (in Russian).
Kiritshenko, A. N., 1951: Nastoyashchie poluzhestkokrylye Evropeyskoy chasti SSSR (Hemiptera). Operedelitel' i bibliografiya. [True bugs of the European part of USSR (Hemiptera). Key and bibliography]. 423 pp. Izdatel'stvo Akademii Nauk SSSR, Moskva & Leningrad.
Linnavuori, R. E., 1953: A Palaearctic Heteropterous material collected by J. Sahlberg and U. Saalas. Annales Entomologici Fennici, 19: 147-167.
Linnavuori, R. E., 1961: Heteroptera of Israel II. Annales Zoologici Societatis Zoologicae Botanicae 'Vanamo', 22: 1-51.
Linnavuori, R. E., 1962: Heteroptera of Israel III. Annales Zoologici Societatis Zoologicae Botanicae 'Vanamo', 24: 1-108.
Linnavuori, R. E., 1999a: Studies on the Miridae fauna of Greece (Hemiptera, Heteroptera). Biologia Gallo-hellenica, 25: 25-68.
Linnavuori, R. E., 1999b: On the genus Phytocoris Fallen (Heteroptera: Miridae, Mirinae) in Iran with remarks on species of the adjacent countries. Part 1. Acta Universitatis Carolinae Biologica, 43: 163-193.
Linnavuori, R. E. & Modarres, M., 1999: Studies on the Heteroptera of the Khorasan province in N.E. Iran. II. Cimicomorpha: Miridae. Entomologica Fennica, 10: 215-231.
Moulet, P., 1995: Faune de France et régiones limitrophes. Vol. 81. Hémiptères Coreoidea (Coreidae, Rhopalidae, Alydidae), Pyrrhocoridae, Stenocephalidae Euro-Méditerranéens. 336 pp. Fédération Française des Sociétés de Sciences Naturelles, Paris.
Péricart, J., 1984: Faune de France et régions limitrophes. Vol. 70. Hémiptères Berytidae Euro-Méditerranéens. 171 pp. Fédération Française des Sociétés de Sciences Naturelles, Paris.
Péricart, J., 1999a. Faune de France et régiones limitrophes. Vol. 84A. Hémiptères Lygaeidae Euro-Méditerranéens. Part 1. xx + 468 pp + 6 pis. Fédération Française des Sociétés de Sciences Naturelles, Paris.
Péricart, J., 1999b: Faune de France et régiones limitrophes. Vol. 84B. Hémiptères Lygaeidae Euro-Méditerranéens. Part 2. iii + 453 pp + 9 pis. Fédération Française des Sociétés de Sciences Naturelles, Paris.
68
©Slovenian Entomological Society, download unter www.biologiezentrum.at
Acta entomologica slovenica, 13 (1), 2005
Josifov, M., 1976: Eine neue Lygaeidae-Art (Heteroptera) aus der Balkanhalbinsel. Acta Zoologica Bulgarica, 4: 65-66.
Josifov, M., 1986: Verzeichnis der von der Balkanhalbinsel bekannten Heteropterenarten (Insecta, Heteroptera). Faunistische Abhandlungen Staatliches Museum für Tierkunde Dresden, 14: 61-93.
Kerzhner, I. M. & Josifov, M., 1999: Miridae Hahn 1833, pp. 1-577. In: Aukema B. & Rieger Ch. (eds.): Catalogue of the Heteroptera of the Palaearctic Region. Vol. 3. Cimicomorpha II. xiv + 577 pp. The Netherlands Entomological Society, Amsterdam.
Kiritshenko, A. N., 1908: K faune Hemiptera - Heteroptera Kryma. [To the Hemiptera - Heteroptera fauna of Crimea]. Russkoe Entomologicheskoe Obozrenie, 8: 234-239 (in Russian).
Kiritshenko, A. N., 1951: Nastoyashchie poluzhestkokrylye Evropeyskoy chasti SSSR (Hemiptera). Operedelitel' i bibliografiya. [True bugs of the European part of USSR (Hemiptera). Key and bibliography]. 423 pp. Izdatel'stvo Akademii Nauk SSSR, Moskva & Leningrad.
Linnavuori, R. E., 1953: A Palaearctic Heteropterous material collected by J. Sahlberg and U. Saalas. Annales Entomologici Fennici, 19: 147-167.
Linnavuori, R. E., 1961: Heteroptera of Israel II. Annales Zoologici Societatis Zoologicae Botanicae 'Vanamo', 22: 1-51.
Linnavuori, R. E., 1962: Heteroptera of Israel III. Annales Zoologici Societatis Zoologicae Botanicae 'Vanamo', 24: 1-108.
Linnavuori, R. E., 1999a: Studies on the Miridae fauna of Greece (Hemiptera, Heteroptera). Biologia Gallo-hellenica, 25: 25-68.
Linnavuori, R. E., 1999b: On the genus Phytocoris Fallen (Heteroptera: Miridae, Mirinae) in Iran with remarks on species of the adjacent countries. Part 1. Acta Universitatis Carolinae Biologica, 43: 163-193.
Linnavuori, R. E. & Modarres, M., 1999: Studies on the Heteroptera of the Khorasan province in N.E. Iran. II. Cimicomorpha: Miridae. Entomologica Fennica, 10: 215-231.
Moulet, P., 1995: Faune de France et régiones limitrophes. Vol. 81. Hémiptères Coreoidea (Coreidae, Rhopalidae, Alydidae), Pyrrhocoridae, Stenocephalidae Euro-Méditerranéens. 336 pp. Fédération Française des Sociétés de Sciences Naturelles, Paris.
Péricart, J., 1984: Faune de France et régions limitrophes. Vol. 70. Hémiptères Berytidae Euro-Méditerranéens. 171 pp. Fédération Française des Sociétés de Sciences Naturelles, Paris.
Péricart, J., 1999a. Faune de France et régiones limitrophes. Vol. 84A. Hémiptères Lygaeidae Euro-Méditerranéens. Part 1. xx + 468 pp + 6 pis. Fédération Française des Sociétés de Sciences Naturelles, Paris.
Péricart, J., 1999b: Faune de France et régiones limitrophes. Vol. 84B. Hémiptères Lygaeidae Euro-Méditerranéens. Part 2. iii + 453 pp + 9 pis. Fédération Française des Sociétés de Sciences Naturelles, Paris.
68
P. Kment, J. Brya, Z. Jindra: New records of true bugs (Heteroptera) of the Balkan peninsula :entri
Péricart, J., 1999c: Faune de France et régiones limitrophes. Vol. 84C. Hémiptères Lygaeidae Euro-Méditerranéens. Part 3. vi + 487 pp + 11 pis. Fédération Française des Sociétés de Sciences Naturelles, Paris.
Péricart, J., 2001a: Family Berytidae Fieber, 1851 - Stilt-bugs, pp. 230-242. In: Aukema B. & Rieger Ch. (eds.): Catalogue of the Heteroptera of the Palaearctic Region. Vol. 4. Pentatomomorpha I. xiv + 346 pp. The Netherlands Entomological Society, Amsterdam.
Péricart, J., 2001b: Family Lygaeidae Schilling, 1829 - Seed-bugs, pp. 35-220. In: Aukema B. & Rieger Ch. (eds.): Catalogue of the Heteroptera of the Palaearctic Region. Vol. 4. Pentatomomorpha I. xiv + 346 pp. The Netherlands Entomological Society, Amsterdam.
Protic, Lj., 1998: Catalogue of the Heteroptera fauna of Yugoslav countries. Part one. Natural History Museum in Belgrade, Special Issue, 38: 1-215.
Protic, Lj., 1999: Biodiversity of the family Berytidae (Heteroptera) in FR Yugoslavia and in the Balkan peninsula. Acta Entomologica Serbica, 4: 11-34.
Protic, Lj., 2001: Catalogue of the Heteroptera fauna of Yugoslav countries. Part two. Natural History Museum in Belgrade, Special Issue, 39: 1-271.
Putshkov, V. G., 1961: Do ekologii malovivchenikh vidiv napivtverdokrilikh Evropeys'koi chastini SRSR. Povidomlennya II. [To the ecology of little known species of Heteroptera of the European part of the USSR. Contribution II]. Pratsi Institutu Zoologii, Akademiya Nauk Ukrains'koi RSR, 17: 86-93 (in Ukrainian).
Putshkov, V. G., 1969: Fauna Ukraini. Tom 21. Vipusk 3. Ligeidi. [Fauna of the Ukraine. Vol. 21/3. Lygaeidae]. 388 pp. Naukova Dumka, Kiiv (in Ukrainian).
Putshkov, V. G. & Putshkov, P. V., 1983: Maloizvestnye poluzhestkokrylye (Heteroptera) yuga SSSR. [Little known bugs (Heteroptera) of the south of the USSR]. Vestnik Zoologii, 1983(3): 17-25 (in Russian, English summary).
Rampazzi, F. & Dethier, M., 1997: Gli Eterotteri (Insecta: Heteroptera) delle torbiere a sfagni del Cantone Ticino e del Moesano (Val Calanca e Val Mescolina - GR), Svizzera. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 70: 419-439.
Rizzotti Vlach, M., di Giovanni, M.V. & Sorice, M., 1996: Sigara basalis (Costa) (Heteroptera, Corixidae), a valid species: diagnostic characters and comparative observations. Italian Journal of Zoology, 63: 261-269.
Seidenstucker, G., 1960: Heteroptera aus Anatolien III. Revue de la Faculté des Sciences de l'Université d'Istanbul, Ser B.: Sciences Naturelles, 25: 145-154.
Seidenstucker, G., 1968: Noch zwei Opisthotaenia-Arten (Heteroptera, Miridae). Reichenbachia, 10: 31-38.
Stehlik, J. L. & Vavrinovâ, I., 1990: Results of the investigation on Hemiptera in Moravia made by the Moravian Museum (Berytidae). Acta Musei Moraviae, Scientiae Naturales, 75: 219-235.
Stys, P., 1976: Faunistic records from Czechoslovakia. Heteroptera: Corixidae: Arctocorisa carinata carinata, Tingidae: Agramma (Serenthiella) minuta,
11
P. Kment, J. Brya, Z. Jindra: New records of true bugs (Heteroptera) of the Balkan peninsula :entri
Péricart, J., 1999c: Faune de France et régiones limitrophes. Vol. 84C. Hémiptères Lygaeidae Euro-Méditerranéens. Part 3. vi + 487 pp + 11 pis. Fédération Française des Sociétés de Sciences Naturelles, Paris.
Péricart, J., 2001a: Family Berytidae Fieber, 1851 - Stilt-bugs, pp. 230-242. In: Aukema B. & Rieger Ch. (eds.): Catalogue of the Heteroptera of the Palaearctic Region. Vol. 4. Pentatomomorpha I. xiv + 346 pp. The Netherlands Entomological Society, Amsterdam.
Péricart, J., 2001b: Family Lygaeidae Schilling, 1829 - Seed-bugs, pp. 35-220. In: Aukema B. & Rieger Ch. (eds.): Catalogue of the Heteroptera of the Palaearctic Region. Vol. 4. Pentatomomorpha I. xiv + 346 pp. The Netherlands Entomological Society, Amsterdam.
Protic, Lj., 1998: Catalogue of the Heteroptera fauna of Yugoslav countries. Part one. Natural History Museum in Belgrade, Special Issue, 38: 1-215.
Protic, Lj., 1999: Biodiversity of the family Berytidae (Heteroptera) in FR Yugoslavia and in the Balkan peninsula. Acta Entomologica Serbica, 4: 11-34.
Protic, Lj., 2001: Catalogue of the Heteroptera fauna of Yugoslav countries. Part two. Natural History Museum in Belgrade, Special Issue, 39: 1-271.
Putshkov, V. G., 1961: Do ekologii malovivchenikh vidiv napivtverdokrilikh Evropeys'koi chastini SRSR. Povidomlennya II. [To the ecology of little known species of Heteroptera of the European part of the USSR. Contribution II]. Pratsi Institutu Zoologii, Akademiya Nauk Ukrains'koi RSR, 17: 86-93 (in Ukrainian).
Putshkov, V. G., 1969: Fauna Ukraini. Tom 21. Vipusk 3. Ligeidi. [Fauna of the Ukraine. Vol. 21/3. Lygaeidae]. 388 pp. Naukova Dumka, Kiiv (in Ukrainian).
Putshkov, V. G. & Putshkov, P. V., 1983: Maloizvestnye poluzhestkokrylye (Heteroptera) yuga SSSR. [Little known bugs (Heteroptera) of the south of the USSR]. Vestnik Zoologii, 1983(3): 17-25 (in Russian, English summary).
Rampazzi, F. & Dethier, M., 1997: Gli Eterotteri (Insecta: Heteroptera) delle torbiere a sfagni del Cantone Ticino e del Moesano (Val Calanca e Val Mescolina - GR), Svizzera. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 70: 419-439.
Rizzotti Vlach, M., di Giovanni, M.V. & Sorice, M., 1996: Sigara basalis (Costa) (Heteroptera, Corixidae), a valid species: diagnostic characters and comparative observations. Italian Journal of Zoology, 63: 261-269.
Seidenstucker, G., 1960: Heteroptera aus Anatolien III. Revue de la Faculté des Sciences de l'Université d'Istanbul, Ser B.: Sciences Naturelles, 25: 145-154.
Seidenstucker, G., 1968: Noch zwei Opisthotaenia-Arten (Heteroptera, Miridae). Reichenbachia, 10: 31-38.
Stehlik, J. L. & Vavrinovâ, I., 1990: Results of the investigation on Hemiptera in Moravia made by the Moravian Museum (Berytidae). Acta Musei Moraviae, Scientiae Naturales, 75: 219-235.
Stys, P., 1976: Faunistic records from Czechoslovakia. Heteroptera: Corixidae: Arctocorisa carinata carinata, Tingidae: Agramma (Serenthiella) minuta,
11
©Slovenian Entomological Society, download unter www.biologiezentrum.at
Acta entomologica slovenica, 13 (1), 2005
Aradidae: Aradus kuthyi, Aradus brevicollis, Aradus brenskei. Acta Entomologica Bohemoslovaca, 73: 61. Wagner, E., 1973: Drei neue Miriden-Arten von Rhodos (Hemipt. Heteropt.).
Nachrichtenblatt der Bayerischen Entomologen, 22: 121-125. Wagner, E., 1975. Die Miridae Hahn, 1831, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Teil 3. Entomologische Abhandlungen, 40(Suppl.): 1-483. Wagner, E. & Weber, H.-H., 1978: Die Miridae Hahn, 1831, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Nachträge zu den Teilen 1-3. Entomologische Abhandlungen, 42(Suppl.): 1-96. Wiprächtiger, P., 2000: Beitrag zur Kenntnis der Wasserwanzenfauna der Ostergauer Weiher in Willisau-Land, Kt. Luzern (Heteroptera: Nepomorpha, Gerromorpha). Entomologische Berichte (Luzern), 43: 11-14. Wröblewski, A., 1980: Pluskwiaki (Heteroptera). Fauna slodkowodna Polski. Zes. 8. [True bugs (Heteroptera). Freshwater fauna of Poland. Vol. 8]. 156 pp. Panstwowe Wydawnictwo Naukowe, Warszawa - Poznan (in Polish).
Received / Prejeto: 10. 3. 2005
52