69
1 ZRC SAZU, Jovan Hadži Institute of Biology, 
Novi trg 2, 1000 Ljubljana, Slovenia
2 University of Ljubljana, Biotechnical Faculty, 
Department of Biology, Večna pot 111, 1000 
Ljubljana, Slovenia
3 Tržaška cesta 62, 1111 Ljubljana
4 University of Maribor, Faculty of Natural 
Sciences and Mathematics, Maribor, Slovenia
* Corresponding author:  
All authors contributed equally to this work.  
For correspondence, see the individual chapters.
Editors:   
Simona Strgulc Krajšek 2 and Tina Klenovšek 4
Citation: Jakob, A., Štampar, L., Lobnik 
Cimerman, Ž., Strgulc Krajšek, S., Vittori, M., 
Dolničar, D., (2024). Significant records of 
plants, algae, fungi, and animals in SE Europe 
and adjacent regions 3. Acta Biologica 
Slovenica 68 (1)
Published: 23.12.2024
https://doi.org/10.14720/abs.68.01.21460
This article is an open access article distributed 
under the terms and conditions of the Creative 
Commons Attribution (CC BY SA) license
Original Research
Significant records  
of plants, algae, fungi,  
and animals in SE Europe  
and adjacent regions, 3
Aljaž Jakob 1, Laura Štampar 2, Žan Lobnik Cimerman 2,  
Simona Strgulc Krajšek 2, Miloš Vittori 2, Dren Dolničar 3
Abstract
In this paper, we present two significant records of bryophytes in Slovenia, 
Cryphaea heteromalla and Riccia cavernosa, which were found in new 
phytogeographical regions. We report the three new records of an isopod 
Armadillo officinalis, a species that has been considered possibly extinct in 
Slovenia, and a new record of bat, Plecotus macrobullaris, from the Julian Alps. 
Keywords
Cryphaea heteromalla; Riccia cavernosa; bryophytes; mosses, liverworts, flora; 
Armadillo officinalis; Isopoda; isopods; Plecotus macrobullaris; Chiroptera; bats; 
fauna; Slovenia
70
Acta Biologica Slovenica, 2025, 68 (1)
Cryphaea heteromalla was hitherto known only from submediterranean parts in the southwestern part of Slovenia (Martinčič 
2024). The localities are in Vipava Valley (Martinčič, 2016), from the area of Škocjan Caves (Strgulc Krajšek et al., 2023) and 
from the flysch Šavrini hills southeast of Koper, where the species is not rare in humid microlocalities (pers. obs.). The new 
record is the first find of this Atlantic-Mediterranean species (Hill et al., 2007) in the central part of Slovenia in the pre-Alpine 
phytogeographical region.
It has been noted this species is benefiting and spreading after the increase in air quality in Europe towards the end of 
20th and beginning of 21st century. However, in Germany, the spread of this species has additionally been linked to climate 
change and eutrophication of the environment (Müller, 2016). Since the new locality is in a highly anthropogenic environ-
ment in intensely managed agricultural land, eutrophication and climate change might likely play a role in the spread of 
this species to continental parts of Slovenia. The species is expected to be found elsewhere in similar anthropogenic and 
eutrophic environments. 
Cryphaea heteromalla is very distinctive when fertile, it has erect sporophyte-bearing shoots with several capsules growing 
along just one side (Figure 1). The capsules are 2.5–3 mm long and have a very short seta (Preston, 2010). In Europe, confu-
sion is only possible with larger Dendrocryphaea lamyana, which grows in regularly flooded habitats, such as river rocks and 
bases of riparian trees, and has blunter leaves (Preston, 2010; Casas, 2006). It is a rare plant with strictly Atlantic-Mediterra-
nean distribution, and it is not expected to occur in Slovenia (Hill et al., 2007).
Funding
This research was funded by Slovenian Research and Innovation Agency (ARIS), grant numbers MR-55879 and P1-0236.
Author(s) Aljaž Jakob
Corresponding author Aljaž Jakob (aljaz.jakob@zrc-sazu.si)
Leg. Aljaž Jakob
Country Slovenia
Statement of significance First record in Slovenia outside the submediterranean region
Locality description Gorenjska, Medvode, Gorenja vas-Reteče, the gravel pit 1.2 km northeast of the village.
Habitat In a partially flooded gravel pit, a semi-permanently flooded riparian forest, on Salix alba bark
Date of observation 2024-06-28
Geographical coordinates N 46.16747309°, E 14.38683831°
Voucher Photo documented
Cryphaea heteromalla (Hedw.) D. Mohr., Crypheaceae (moss, bryophyte)
71
Acta Biologica Slovenica, 2025, 68 (1)
Riccia cavernosa (Hoffm.) Hoffm. is a temperate species of a complex thalloid liverwort with a circumpolar distribution. It is 
also present in the tropics and the Southern Hemisphere (Hill & Preston, 1998). It is typically found in temporarily wet habitats 
such as the banks of ponds, ditches, rivers, wetlands, and other periodically inundated sites (Frey et al., 2006). 
R. cavernosa is a species from subgenus subgenus Ricciella (A. Braun) Boulay, section Spongodes Nees (Hodgetts et al., 
2020). It is characterised by very distinctly perforated, yellow-green thallus without reddish colouration. A short, inconspicu-
ous groove is visible on the upper surface only at the tip of the lobes (Long, 2010). The species has significant morphological 
similarities to R. crystallina L., therefore ripe spores are needed for reliable identification (Paton, 1999).
Author(s) Laura Štampar, Žan Lobnik Cimerman, Simona Strgulc Krajšek
Corresponding author Simona Strgulc Krajšek (simona.strgulc@bf.uni-lj.si)
Leg. Laura Štampar
Country Slovenia
Statement of significance The third known locality in Slovenia, first in the dinaric phytogeographical region
Locality description Notranjska, Cerkniško Jezero, Laze pri Gornjem Jezeru, left bank of river Jezerščica, next to the bridge near the Malnšče observation point, 550 m a.s.l.
Habitat damp sandy soil
Date of observation 2024-08-30
Geographical coordinates N 45.728611°, E 14.404167° 
Voucher Herbarium LJU (s.n.)
Riccia cavernosa (Hoffm.) Hoffm., fam. Ricciaceae (liverwort, bryophyte)
Figure 1. Cryphaea heteromala with characteristic erect shoots with 
several laterally growing capsules (photo: A. Jakob).
72
Acta Biologica Slovenica, 2025, 68 (1)
R. cavernosa is listed as the least concerned (LC) in the European Red List (Hodgetts et al., 2019). It is widespread in Europe 
but rare in the Mediterranean region (Blockeel et al., 2014), and its conservation status varies significantly across different 
countries. In Italy and Austria, it is endangered (EN), in Hungary, it is near threatened (NT), and in Croatia, it has no conser-
vation status (Hodgetts and Lockhart, 2020). In Slovenia, it was found for the first time in 2021 in the vicinity of Maribor by Ž. 
Lobnik Cimerman (Ellis et al., 2022) and later also near Dokleževje in the Prekmurje Region (Lobnik Cimerman et al., 2023). 
In Slovenia, it has a status of endangered species (Martinčič, 2024). 
Our sample was collected from a sandy substrate on the left bank of the Jezerščica River, approximately 80 meters down-
stream from the bridge near the Malnšče observation point. The Jezerščica River is part of the Stržen River, flowing near 
Gorenje Jezero, in the vicinity of intermittent Cerknica Lake. The Stržen is one of the main streams of the lake, flowing 
through this karst region and frequently forming seasonal floodplains. The banks of the Stržen River host numerous other 
plant species, mainly amphibious macrophytes, due to the constant water level fluctuation, creating a dynamic environment 
ideal for their growth. Some of the species of amphibious macrophytes with the highest abundance, which had already been 
recorded by Kržič et al. (2007), and were confirmed by our observations, are Myosotis scorpioides, Nuphar luteum, Ranun-
culus trichophyllus, Rorippa amphibia and Schoenoplectus lacustris.
Riccia cavernosa appeared in considerable abundance at the locality, covering entire sections of the riverbank (Figure 2), 
which is characteristic of the species when water level drops and its ecological requirements are met. Some field obser-
vations suggest that its spores can remain viable for several years (Blockeel et al., 2014). It appears that the current condi-
tions are favourable for its growth, as the species is well adapted to fluctuations in water levels, typical of the area around 
Cerknica Lake and the Stržen River. However, excessive overgrowth of the riverbank could potentially threaten R. cavernosa.
Funding
This work was supported by the Slovenian Research Agency (ARIS) through the programme group P1-0212 and ARIS Young 
research grant (Ž. Lobnik Cimerman).
Figure 2. Riccia cavernosa growing on humid sandy soil 
(photo: L. Štampar).
73
Acta Biologica Slovenica, 2025, 68 (1)
The pill bug Armadillo officinalis Duméril, 1816 was first recorded on the territory of Slovenia in Škocjan near Koper in July 
1926 (Strouhal, 1929). This record was the basis for the inclusion of the species in checklists of the Slovenian terrestrial 
isopod fauna (Potočnik, 1989; Vittori et al., 2023), but the species was not recorded in Slovenia again for nearly a century. 
Investigations of the terrestrial isopod fauna of the Slovenian coast in the 1970s and 1980s did not detect A. officinalis 
(Potočnik, 1984). As a result, the species was first considered endangered (Potočnik, 1992), and later, as efforts to find it had 
failed, possibly extinct in Slovenia (Potočnik, 1996). Despite this, it was not assigned a red list status in the national Red list 
of Malacostraca (Ur. l. RS., 2002).
The first recent observation was of a single isopod under a wooden board at the saltpans at Fontanigge in September 2023 
(Figure 3A). Two days later, more than ten individuals were observed near the Walls of Piran, again under a wooden board. 
Six isopods found there were collected and preserved for further examination (Figure 3B). A third observation occurred at 
Fontanigge in March 2024. This time, more than ten individuals were observed under wooden pallets near the Museum of 
Salt Making (Figure 3C).
Species of the genus Armadillo are capable of rolling into a ball, with the flat protopodites of the uropods completing the 
outline of the pleon, and the uropod exopodites reduced to small cones that insert dorsally (Figure 3B). The genus is further 
characterized by monospiracular lungs on all pleopod exopodites and by modifications of pereopods 4 and 5 that enable 
these isopods to stridulate (Schmalfuss, 1996; Taiti et al., 1998; Cividini et al., 2020). While several species of Armadillo have 
been described from the Eastern Mediterranean, North Africa, and the Iberian Peninsula, A. officinalis is the only widespread 
Mediterranean species (Schmalfuss, 2003). It can be distinguished on the basis of the shape of the uropod protopodites and 
the pleotelson, the smoothness of the tergites, and male characters, mainly the shape of pereopod 7 and the exopodite of 
pleopod 1 (Schmalfuss, 1996).
Author(s) Miloš Vittori
Corresponding author Miloš Vittori (milos.vittori@bf.uni-lj.si) 
Leg. (1,2) Miloš Vittori(3) Barbara Breznik, Barbora Ďurajková, Miloš Vittori 
Country Slovenia
Statement of significance New records of a species considered possibly extinct in Slovenia
Locality description
(1) Sečovlje, Fontanigge, elevation 2 m.a.s.l.
(2) Piran, by the Walls of Piran, elevation 35 m.a.s.l.
(3) Sečovlje, Fontanigge, by the Museum of Salt Making, elevation 2 m.a.s.l.
Habitat
(1) Mediterranean halophilous scrub
(2) Paved street
(3) Mediterranean halophilous scrub
Date of observation
(1) 2023-09-21
(2) 2023-09-23
(3) 2024-03-30
Geographical coordinates
(1) N 45.47202°, E 13.6034°
(2) N 45.52688°, E 13.57058°
(3) N 45.47636°, E 13.59347°
Voucher
Six specimens collected in Piran are kept in the isopod collection of Miloš Vittori, University of 
Ljubljana, Biotechnical Faculty, Department of Biology, Večna pot 111, 1000 Ljubljana, Slovenia. 
Individuals observed at Fontanigge were not collected and were photographed instead.
Armadillo officinalis Duméril, 1816, fam. Armadillidae (animal)
74
Acta Biologica Slovenica, 2025, 68 (1)
The consistent presence of these isopods suggests that the population of A. officinalis in Slovenia may be recovering, with 
climate change possibly contributing to its success. Sket (2003) considered the species to be expanding its range, aided by 
human activity. Nevertheless, its recorded distribution in the country remains limited to the immediate proximity of the coast.
Acknowledgement
This work was supported by University Infrastructural Centre “Microscopy of Biological Samples” at the Biotechnical Faculty, 
University of Ljubljana.
Funding
This research received no external funding.
Figure 3. Armadillo officinalis. A: The individual 
observed at Fontanigge in September 2023. B: One of 
the specimens collected in Piran in September 2023. 
The uropod protopodites complete the outline of the 
pleon and the uropod exopodites are diminutive (arrow-
head). C: Individuals photographed at Fontanigge in 
March 2024. (photos A, B, C: Miloš Vittori).
75
Acta Biologica Slovenica, 2025, 68 (1)
On 11 July 2024, I found 2 individuals of Plecotus in the space between the planks and beams of the barn (WGS 84: 46.287972° 
N, 13.763278° E; Figure 4) next to the mountain hut Koča pod Bogatinom (address: Ukanc 148), on the Komna plateau, at 
1515 m a.s.l. One was too far away to determine the species, while the other one, which was closer, I was able to identify as 
Plecotus macrobullaris based on the characteristic inverted triangular shape of the lower lip (Dietz & Kiefer, 2016). The bat, 
identified as P. macrobullaris, was initially positioned much closer to the edge of the gap between the barn’s planks (Figure 
4B), with its dark greyish triangular lower lip (Dietz & Kiefer, 2016) clearly visible. However, it retreated deeper into the gap 
after we disturbed it with light while attempting to photograph it.
The record is the highest roost of this species in Slovenia, since the previously highest elevation known roost was in the 
church sv. Duha in Podolševa at 1250 m a.s.l. (Presetnik et al., 2023). It is the third highest observation overall, as the highest 
published record originates from the mist netting at Planina pri Jezeru at 1750 m a.s.l. (Alberdi et al., 2013; CKFF, 2024) and 
the second highest is an ultrasound recording from Planina Krstenica at 1670 m a.s.l. (A. Zamolo, personal communication, 
August 28, 2024). According to the general species vertical distribution, the finding of P. macrobularis at this altitude on 
Komna is not surprising. The species is common in the Alpine regions and is found in all the main mountain ranges of the 
western Palearctic, stretching from the Pyrenees to the Middle East, and is distributed at altitudes ranging from sea level 
to 2800 m a.s.l. (Alberdi et al., 2013). The scarcity of observations of P. macrobullaris in Slovene montane belt (CKFF, 2024) 
therefore indicates a lack of research in the Slovenian high mountains.
Acknowledgement
I would like to thank Primož Presetnik for gathering and sharing the relevant data about P. macrobullaris in Slovenia and for 
reviewing the first draft of this article and Aja Zamolo for sharing her records of this species. I would also like to thank Simona 
Strgulc Krajšek for sharing her photograph of P. macrobullaris that she took when I found it in the barn.
Funding
This research received no external funding.
Author(s) Dren Dolničar
Corresponding author Dren Dolničar (dren.dolnicar@gmail.com)
Leg. Dren Dolničar
Country Slovenia
Statement of significance The highest elevation of recorded roost site for this species in Slovenia.
Locality description Triglav National Park, Komna plateau, 1515 m a.s.l.
Habitat An alpine meadow below the tree line, with a roost site in a wooden barn.
Date of observation 2024-07-11
Geographical coordinates N 46.287972°, E 13.763278° 
Voucher /
Plecotus macrobullaris Kuzyakin, 1965 (Chiroptera, Mammalia)  
(bats, mammals)
76
Acta Biologica Slovenica, 2025, 68 (1)
Figure 2. A) Plecotus macrobullaris; B) the roosting place; C) the barn near the hut Koča pod Bogatinom on Komna, where at least two Ple-
cotus bats were roosting (photo: A: Simona Strgulc Krajšek; B, C: Dren Dolničar).
A B C
References
Alberdi, A., Garin, I., Aizpurua, O., Aihartza, J., 2013. Review on the Geographic and Elevational Distribution of the Mountain Long-Eared Bat Plecotus macrobullaris, 
Completed by Utilising a Specific Mist-Netting Technique. Acta Chiropterologica 15(2): 451–461. https://doi.org/10.3161/150811013X679071. 
Blockeel, T. L., Bosanquet, S. D. S., Hill, M. O., Preston, C. D. (eds.), 2014. Atlas of British and Irish bryophytes: the distribution and habitat of mosses and liverworts 
in Britain and Ireland. Pisces Publications on behalf of the British Bryological Society.
Casas, C., Brugués, M., Cros, R. M., & Sérgio, C., 2006. Handbook of Mosses of the Iberian Peninsula. p. 311.
Cividini, S., Sfenthourakis, S., Montesanto, G., 2020. Are terrestrial isopods able to use stridulation and vibrational communication as forms of intra and 
interspecific signaling and defense strategies as insects do? A preliminary study in Armadillo officinalis. The Science of Nature, 107(1), 4. https://doi.org/10.1007/
s00114-019-1656-3.
CKFF, 2024. Plecotus macrobullaris. Database of Center za kartografijo favne in flore, Miklavž na Dravskem polju. (July 2024, prepared by Primož Presetnik).
Dietz, C., Kiefer, A., 2016. Bats of Britain and Europe. Bloomsbury USA.
Ellis, L. T., Afonina, O. M., Czernyadjeva, I. V., Alegro, A., Šegota, V., Boiko, M., Zagorodniuk, N., Burghardt, M., Alataş, M., Aslan, G., Batan, N., Dragićević, S., Erata, 
H., Kırmacı, M., Özenoğlu, H., Evangelista, M., Valente, E. B., Feletti, T. A., Ezer, T., Fedosov, V. E., Fuertes, E., Oliván, G., Natcheva, R., Gospodinov, G., Hodgson, 
A., Kiebacher, T., Köckinger, H., von Konrat, M., Strgulc Krajšek, S., L. Cimerman, Ž., Kučera, J., Mikulášková, E., Müller, F., Muñoz, J., Ochyra, R., Peralta, D. F., 
Philippe, M., Porley, R. D., Rawat, K. K., Paul, R. R., Ros, R. Werner, M., O., Schäfer-Verwimp, A., Sérgio, C., Shkurko, A. V., Söderström, L., de Souza, A. M., Spitale, 
D., Ștefănuţ, S., Tabua, M., Winter, G., 2022. New national and regional bryophyte records, 69, Journal of Bryology, 44:1, 87–102. https://doi.org/10.1080/03736
687.2022.2061242. 
Frey, W., Frahm, J. P., Fischer, E., Lobin, W., Blockeel, T. L. (eds.), 2006. The Liverworts, Mosses and Ferns of Europe. Harley Books, Essex, 512 pp.
Hill, M. O., Preston, C. D., 1998, The geographical relationships of British and Irish bryophytes, Journal of Bryology, 20:1, 127–226, https://doi.org/10.1179/
jbr.1998.20.1.127. 
Hill, M. O., Preston, C. D., Bosanquet, S. D. S., & Roy, D. B., 2007. BRYOATT : attributes of British and Irish mosses, liverworts and hornworts. Centre for Ecology 
and Hydrology.
Hodgetts, N. G., Lockhart, N., 2020. Checklist and country status of European bryophytes – update 2020. – Irish Wildlife Manuals 123. – Dublin: Department of 
Culture, Heritage and the Gaeltacht, 224 pp.
Hodgetts, N. G., Söderström, L., Blockeel, T. L., Caspari, S., Ignatov, M. S., Konstantinova, N. A., Lockhart, N., Papp, B., Schröck, C., Sim-Sim, M., Bell, D., Bell, 
N. E., Blom, H. H., Bruggeman-Nannenga, M. A., Brugués, M., Enroth, J., Flatberg, K. I., Garilleti, R., Hedenäs, L., Holyoak, D. T., Hugonnot, V., Kariyawasam, I., 
Köckinger, H., Kučera, J., Lara, F., Porley, R. D., 2020. An annotated checklist of bryophytes of Europe, Macaronesia and Cyprus. Journal of Bryology 42: 1–116.
77
Acta Biologica Slovenica, 2025, 68 (1)
Hodgetts, N., Cálix, M., Englefield, E., Fettes, N., García Criado, M., Patin, L., Nieto, A., Bergamini, A., Bisang, I., Baisheva, E., Campisi, P., Cogoni, A., Hallingbäck, 
T., Konstantinova, N., Lockhart, N., Sabovljević, M., Schnyder, N., Schröck, C., Sérgio, C., Sim Sim, M., Vrba, J., Ferreira, C. C., Afonina, O., Blockeel, T., Blom, H., 
Caspari, S., Gabriel, R., Garcia, C., Garilleti, R., González Mancebo, J., Goldberg, I., Hedenäs, L., Holyoak, D., Hugonnot, V., Huttunen, S., Ignatov, M., Ignatova, 
E., Infante, M., Juutinen, R., Kiebacher, T., Köckinger, H., Kučera, J., Lönnell, N., Lüth, M., Martins, A., Maslovsky, O., Papp, B., Porley, R., Rothero, G., Söderström, 
L., Ştefănuţ, S., Syrjänen, K., Untereiner, A., Váňa, J., Vanderpoorten, A., Vellak, K., Aleffi, M., Bates, J., Bell, N., Brugués, M., Cronberg, N., Denyer, J., Duckett, J., 
During, H. J., Enroth, J., Fedosov, V., Flatberg, K. I., Ganeva, A., Gorski, P., Gunnarsson, U., Hassel, K., Hespanhol, H., Hill, M., Hodd, R., Hylander, K., Ingerpuu, N., 
Laaka-Lindberg, S., Lara, F., Mazimpaka, V., Mežaka, A., Müller, F., Orgaz, J. D., Patino, J., Pilkington, S., Puche, F., Ros, R. M., Rumsey, F., Segarra-Moragues, J. 
G., Seneca, A., Stebel, A., Virtanen, R., Weibull, H., Wilbraham, J., Żarnowiec, J., 2019. A miniature world in decline: European Red List of Mosses, Liverworts and 
Hornworts. IUCN, Brussels. https://doi.org/10.2305/IUCN.CH.2019.ERL.2.en. 
Lobnik Cimerman, Ž, Kopitar, D, Strgulc Krajšek, S., 2023. Arable bryophytes from Northeastern Slovenia with new and interesting national records. Botanica 
Serbica 47(2): 301–308. https://doi.org/10.2298/BOTSERB2302301C. 
Long, D., 2010. Riccia cavernosa, Cavernous Crystalwort. In: Atherton, I., Bosanquet, S., Lewley, M. (eds.), Mosses and Liverworts of Britain and Ireland, a field 
guide. British Bryological Society, p 263.
Martinčič, A., 2016. Updated Red List of bryophytes of Slovenia. Hacquetia, 15(1), 107–126. https://doi.org/10.1515/hacq-2016-0006.
Martinčič, A., 2024. New checklist and the Red list of the hornworts (Anthocerotophyta) and liverworts (Marchantiophyta) of Slovenia. Hacquetia 23 (2): 175–197. 
https://doi.org/10.2478/hacq-2023-0013. 
Martinčič, A., 2024. New Checklist and the Red list of the mosses (Bryophyta) of Slovenia. Hacquetia, 23(1), 69–118. https://doi.org/10.2478/hacq-2023-0006.
Müller, J., 2016. Einwanderung von Cryphaea heteromalla (Hedw.) D. Mohr nach Berlin und Brandenburg. Veröffentlichungen Des Naturkundemuseums 
Potsdam 2: 37–43.
Paton, A. J., 1999. The liverwort flora of the British Isles. Colchester: Harley Books.
Potočnik, F., 1984. Mokrice (Isopoda terrestria) slovenske obale Jadrana. Biološki vestnik, 32(1), 77–86.
Potočnik, F., 1989. The survey of the fauna of woodlices (Isopoda terrestria) of Yugoslavia. Biološki vestnik, 37, 61–82.
Potočnik, F., 1992. The red list of endangered Isopoda terrestria (Crustacea) in Slovenia. Varstvo narave, 17, 157–164.
Potočnik, F., 1996. Mokrice (Isopoda Terrestria). In: Gregori J, Martinčič A, Tarman K, Urbanc-Berčič O, Tome D, Zupančič M (eds.): Narava Slovenije, stanje in 
perspektive. Društvo ekologov Slovenije, Ljubljana, pp. 247–251.
Presetnik, P., Zamolo, A., Grgurevič, S., Šalamun, A., 2023. Monitoring populacij izbranih ciljnih vrst netopirjev v letih 2021–2023. Končno poročilo. Center za 
kartografijo favne in flore, Miklavž na Dravskem polju. 212 str., digitalne priloge. [Naročnik: Ministrstvo za naravne vire in prostor, Ljubljana].
Preston, C., 2010. Cryphaea heteromalla, Lateral Cryphaea. In: Atherton, I., Bosanquet, S., Lewley, M. (eds.), Mosses and Liverworts of Britain and Ireland, a field 
guide. British Bryological Society, p 672.
Schmalfuss, H., 1996. The terrestrial isopod genus Armadillo in Western Asia (Oniscidea: Armadillidae), with descriptions of five new species. Stuttgarter 
Beiträge zur Naturkunde, Serie A, 544.
Schmalfuss, H., 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beiträge zur Naturkunde, Serie A, 654.
Sket, B., 2003. Raki – Crustacea. In: Sket B, Gogala M, Kuštor V (eds.): Živalstvo Slovenije. Tehniška založba Slovenije, Ljubljana, pp. 188–224.
Šraj-Kržič, N., Germ, M., Urbanc-Berčič, O., Kuhar, U., Janauer, G. A., Gaberščik, A., 2007. The Quality of the Aquatic Environment and Macrophytes of Karstic 
Watercourses. Plant Ecology 192(1): 107–118. 
Strgulc Krajšek, S., Gey, S., Cezanne, R., Eichler, M., 2024. Report on the annual excursion of BLAM to Slovenia from September 13th to 17th, 2023. Herzogiella 
11: 13–26. 
Strouhal, H., 1929. Die Landisopoden des Balkans. 3. Beitrag: Südbalkan. Zeitschrift für wissenschaftliche Zoologie, 133, 57–120.
Taiti, S., Paoli, P., Ferrara, F., 1998. Morphology, biogeography, and ecology of the family Armadillidae (Crustacea, Oniscidea). Israel Journal of Zoology, 44, 
291–301.
Ur. l. RS., 2002. Pravilnik o uvrstitvi ogroženih rastlinskih in živalskih vrst v rdeči seznam. Uradni list RS, no. 82/02, 42/10.
Vittori, M., Mavrič, B., Pekolj, A., Govedič, M., Zagmajster, M., 2023. A checklist of isopods (Crustacea: Isopoda) in Slovenia. Natura Sloveniae, 25(2), 27–52. 
https://doi.org/10.14720/ns.25.2.27-52.