ANNALES • Ser. hist. nat. • 14 • 2004 • 2
review article	UDC 597.5:591.134(262-18)
received: 2004-11-14
LENGTH-WEIGHT RELATIONSHIPS OF FISHES FROM TURKISH SEAS:
A REVIEW
Hatice TORCU KO< & Zeliha AKA ERDOGAN University of Balikesir, Faculty of Science and Arts, Department of Biology, TR-10100 Balikesir, Turkey
jakov DULCIC
Institute of Oceanography and Fisheries, HR-21000 Split, P.O. BOX 500, Croatia E-mail: dulcic@izor.hr
ABSTRACT
The paper presents 360 length-weight relationships gathered from available literature pertaining to 90 fish species belonging to 40 families occurring throughout Turkish seas. The value of the slope b ranged from 0.409 for Thunnus thynnus to 4.343 for Dentex macrophthalmus. The mean value of b was 3.088 (± SD = 0.898) and did not differ significantly from 3 (t-test, p <0.05). The median value of b was 3.062 and 50% of b values ranged between 2.941 and 3.190.
Key words: length-weight relationship, fishes, Turkish seas, review
RAPPORTO PESO/LUNGHEZZA DEI PESCI NEI MARI DELLA TURCHIA: UNA RASSEGNA
SINTESI
Sulla base dei dati messi a disposizione dall'attuale letteratura, l'articolo elenca 360 esempi di rapporto peso/lunghezza di pesci appartenenti a 90 specie e 40 famiglie che dimorano nei mari della Turchia. La curva (b) va dallo 0,409 della specie Thunnus thynnus al 4,343 di quella del Dentex macrophthalmus. Il valore medio di b e 3,088 (± SD = 0,898) e si discosta pochissimo dal 3 (t-test, p<0,05). La mediana del coefficente b e di 3,062, mentre il 50 % del valore di b oscilla tra 2,941 e 3,190.
Parole chiave: rapporto peso/lunghezza, pesci, mari della Turchia, rassegna
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INTRODUCTION
Recording length and weight data is a standard procedure of fish sampling programs. Such data are essential for studies of fish population dynamics. Length-weight relationships have a number of uses including: (a) the estimation of standing-crop biomass when the length-frequency distribution is known; (b) estimation of biomass from length observations because of technical difficulties (e.g. bobbing motion of the boat) and the amount of time required to record weight in the field; (c) the calculation of condition indexes in the analysis of ontogenetic changes; and (d) comparing the life stories and morphological variations of fish populations from different regions (Cailliet et a/.,1986; Safran, 1992; Pe-trakis & Stergiou, 1995; Goncalves et a/., 1996; MartinSmith, 1996; Morato et a/., 2001; Stergiou & Motopou-los, 2001; Filiz & Bilge, 2004).
In this review, we gathered 336 length-weight relationships from the available literature for 70 fish species from Turkish seas.
MATERIAL AND METHODS
All length-weight relationships presented here are the product of field studies conducted during 1972-2004 in Turkish seas. For the majority of the original length-weight relationship (W=aLb), length was expressed in cm and weight in g (356 relationships out of 360; 98.8%), whereas for 4 (1.2%) relationships, length and weight were expressed in mm and g.
RESULTS AND DISCUSSION
Overall, 360 length-weight relationships, gathered from the literature, represent 90 species and 40 families occurring in Turkish marine waters (Tab. 1). Overall, 50 out of the 360 length-weight relationships refer to Mer-/uccius mer/uccius (14.9%), 35 (10.4%) to Mu//us bar-batus, and 26 (7.7%) to Page//us erythrinus, three of the most commercially important demersal species in Turkish seas. Regarding the number of studies, of which length-weight relationship of an individual family is recorded, Sparidae dominated the overall records (84, 25.0%), and followed by Mullidae (60, 17.9%), Merluc-cidae (50, 14.9%), Engraulidae (17, 5.1%) and Mugili-dae (14, 4.2%). Each of the remaining 28 families was represented by less than one relationship.
The value of the slope b in the plot of log W against log W ranged from 0.409 for Thunnus thynnus in Aegean and Mediterranean Sea to 4.343 for Dentex mac-
rophthalmus in Saros Bay. The mean value of b was 3.088 (± SD = 0.8985) and did not differ significantly from 3 (t-test, p <0.05). The median value of b was 3.062 and 50 % of b values ranged between 2.941 and 3.19.
According to Stergiou & Moutopoulos (2001), a plot of a versus b for all known length-weight relationships of a species results in a linear relationship, and this relationship can be used to identify outliers. We have applied this method to all species with more than 10 length-weight relationships. This led to the detection of outliers, where the respective point deviated more than two standard deviations from the regression line. These length-weight relationships were marked as "questionable" in Table 1.
The length-weight relationship in fishes is affected by a number of factors, including season, habitat, gonad maturity, sex, diet and stomach fullness, health and preservation techniques (Dulčič & Kraljevič, 1996), all of which were not accounted for in this review. As stressed by Bagenal & Tesch (1978) and Petrakis & Stergiou (1995), the use of relationships should be limited to the size range used to estimate the parameters. Extrapolation to juvenile or immature stages may be particularly unwise given the potential differences in growth patterns between adults and earlier life history stages (Bagenal & Tesch, 1978). The parameters of the length-weight relationship may vary significantly according to season, habitat and even on a daily basis (ibid). In fishes, size is generally more biologically relevant than age, mainly because several ecological and physiological factors are more size-dependent than age-dependent (Dulčič & Kraljevič, 1996). Consequently, variability in size has important implications for diverse aspects of fisheries science and population dynamics. Length-weight relationships have several applications, namely on fish biology, physiology, ecology and fisheries assessment. In biological studies, length-weight relationships enable seasonal variations in fish growth to be followed and the calcultaion of condition indexes. In fisheries studies, length-weight relationships have many different uses, including the estimation of weight from length, the estimation of weight-at-age and the conversion of growth-in-length equations to growth-in-weight. The establishment of length-weight relationships is also fundamental for the calculation of production and biomass of a fish population, being very useful for biomass estimations based on visual census of fish populations.
This review of length-weight parameters reported here will be of considerable use in ongoing studies of the catch in Turkish commercial fisheries.
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Tab. 1: Parameters of the length-weight relationship W = aLb of fish species from Turkish marine waters (length (cm), weight (g)).
Legend: Sex (M, male; F, female; C, combined; U, unidentified); Year = year of sampling; S = sampling season (AUT, Autumn; Wl, Winter; SP, Spring; SU, Summer; C, All seasons); L = length (TL, Total length; FL, Fork length; SL, Standard length); a = intercept (from the literature); b = slope (of the relationship W=aLb); r2 = coefficient of correlation; N = sampling size; min and max = minimum and maximum sample lengths (cm); SD = standard deviation.
Species are listed in alphabetical order.
Tab. 1: Parametri razmerja W = aLb med dolžino in težo ribjih vrst, živečih v turških morjih (dolžina (cm), teža (g)).
Legenda: Sex = spol (M, samec; F, samica; C, oba skupaj; U, neidentificiran); Year = leto vzorčenja; S = sezona vzorčenja (AUT, jesen; Wl, zima; SP, pomlad; SU, poletje; C, vse sezone); L = dolžina (TL, celotna dolžina; FL, dolžina razcepa repa; SL, standardna dolžina); a = presečišče (na podlagi virov); b = naklon (iz razmerja W=aLb); r2 = korelacijski koeficient; N = število primerkov; min in max = najmanjše in največje vzorčevalne velikosti (cm); SD = standardni odklon. Vrste so navedene v abecednem redu.
Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Alosa sp.	Black Sea	C	1988-94	C	TL	0.0081	3.103	0.921*	65					Erkoyuncu et al. (1994)
Arnoglossus laterna	Izmir Bay	C	1996-97	C	TL	0.00932	2.897	0.940	643	12.1	6.0	15.3		Mater & Bayhan (2000)
Argentina sphyraena	Sigacik trawl area	C	2003	C	TL	0.0062	2.93	0.93	238	-	7.5	20.7	-	Filiz & Bilge (2004)
Atherina hepsetus	Eceabat Dardanelles	C	1997-99	C	FL	0.01287	2.890	0.999	564	-	5.8	13.0	-	Altun (2000)
Belone belone	Black Sea	C	1988-94	C	TL	0.0005	3.203	0.940*	65					Erkoyuncu et al. (1994)
Belone belone euxini	Black Sea	C	1995-96	C	TL	0.00055	3.177	0.940*	643	37.55	31.2	52.2	0.17	Samsun (1995)
Capros aper	Yejilova Bay	F	1992	SU	TL	0.0000294	2.912	0.883*	170	86.35			0.7	Kaya & Ozaydin (1996)
Capros aper	Yejilova Bay	M	1992	SU	TL	0.0000204	2.915	0.883*	165	84.72			0.6	Kaya & Ozaydin (1996)
Capros aper	Sigacik trawl area	C	2003	Wl, SU	TL	0.0232	2.83	0.98	455		2.9	10.1		Filiz & Bilge (2004)
Cepola rubescens	Izmir Bay	F	1996-97	C	TL	0.32388	1.269	0.792	131	34.43	11.5	45.6	6.76	Kaya et al. (2001)
Caelorinchus caelorinchus	Sigacik trawl area	C	2003	C	TL	0.0065	2.74	0.78	208	-	4.0	21.6	-	Filiz & Bilge (2004)
Cepola rubescens	Izmir Bay	M	1996-97	C	TL	0.2154	1.384	0.810	144	29.17	19.8	47.1	6.26	Kaya et al. (2001)
Chelidonichthyes lucerna	Iskenderun Bay	F	1999-00	C	TL	0.0095	2.990	0.960*	199		8.0	30.3		Ismen et al. (2004)
Chelidonichthyes lucerna	Iskenderun Bay	M	1999-00	C	TL	0.0089	3.010	0.980*	143		8.3	21.2		Ismen et al. (2004)
Chelon labrosus	Gulluk Lagoon	C	1993-94	C	TL	0.0075539	3.067	0.962*	45	26.4	20.0	35.0	3.5	Hossucu (2001)
Chlorophthalmus agassizi	Sigacik trawl area	C	2003	C	TL	0.0027	3.37	0.98	378	-	7.7	17.5	-	Filiz & Bilge (2004)
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Chimaera	Sigacik trawl	C	2003	C	TL	0.0028	2.82	0.98	17	-	13.1	75.3	-	Filiz & Bilge
monstrosa	area													(2004)
Conger conger	Sigacik trawl area	C	2003	C	TL	0.0005	3.24	0.96	22	-	32.2	65.4	-	Filiz & Bilge (2004)
Dasyatis	iskenderun	F	1999-00	C	TL	0.00091	3.440	0.940	110	-	20.5	88.0	-	Ismen
pastinaca	Bay													(2003)
Dasyatis	iskenderun	M	1999-00	C	TL	0.00237	3.170	0.950	146	-	20.0	73.0	-	Ismen
pastinaca	Bay													(2003)
Dasyatis	iskenderun	C	1999-00	C	TL	0.00144	3.310	0.940	256	-	20.0	88.0	-	Ismen
pastinaca	Bay													(2003)
Dasyatis	Sigacik trawl	C	2003	Wl,	TL	0.0149	2.81	0.85	29	-	37.3	74.2	-	Filiz & Bilge
pastinaca	area			su										(2004)
Dentex dentex	iskenderun Bay	C	2000	AuT	TL	0.0861	2.495	0.921	16	42.41	31.40	51.0	6.35	Can et al. (2000)
Dentex gibbosus	iskenderun	C	2000	AuT	TL	0.0341	2.714	0.857	34	27.29	17.68	47.3	6.83	Can et al.
	Bay													(2000)
Dentex	Saros Bay	C	1991	su	TL	0.01760	3.071	0.864*	20	-	-	-	-	Benli et al.
macrophthalmus														(2000)
Dentex	izmir Bay	C	1991	su	TL	0.02248	2.966	0.889*	20	-	-	-	-	Benli et al.
macrophthalmus														(2000)
Dentex	Gulluk Bay	C	1991	su	TL	0.00782	3.292	0.974*	20	-	-	-	-	Benli et al.
macrophthalmus														(2000)
Dentex	Gokova Bay	C	1991	su	TL	0.02404	2.932	0.874*	60	-	-	-	-	Benli et al.
macrophthalmus														(2000)
Dentex	Saros Bay	C	1992	Wl	TL	0.01974	3.027	0.988*	12	-	-	-	-	Benli et al.
macrophthalmus														(2000)
Dentex	Saros Bay	C	1992	sp	TL	0.00002	4.343	0.980*	58	-	-	-	-	Benli et al.
macrophthalmus														(2000)
Dentex	Gokova Bay	C	1992	sp	TL	0.01280	3.107	0.970*	40	-	-	-	-	Benli et al.
macrophthalmus														(2000)
Dentex	Gokova Bay	C	1993	sp	TL	0.01731	3.084	0986*	46	-	-	-	-	Benli et al.
macrophthalmus														(2000)
Dentex	izmir Bay	C	1993	aut	TL	0.02968	2.859	0.975*	9	-	-	-	-	Benli et al.
macrophthalmus														(2000)
Dentex	Gulluk Bay	C	1993	aut	TL	0.02946	2.812	0.974*	10	-	-	-	-	Benli et al.
macrophthalmus														(2000)
Dentex	Gokova Bay	C	1993	aut	TL	0.02117	2.977	0.978*	82	-	-	-	-	Benli et al.
macrophthalmus														(2000)
Dentex	Aegean Sea	F	1991-93	C	FL	0.01308	3.171	0.955*	285	14.10	8.2	22.0	0.18	Ozaydin
macrophthalmus														(1997)
Dentex	Aegean Sea	M	1991-93	C	FL	0.00921	3.282	0.941*	203	14.65	8.2	22.0	0.37	Ozaydin
macrophthalmus														(1997)
Dentex	Aegean Sea	C	1991-93	C	FL	0.01191	3.198	0.941*	507	14.23	7.8	22.0	0.20	Ozaydin
macrophthalmus														(1997)
Dentex	Marmara,	C	1991-93	Wl	FL	0.0000099	3.143	0.962*	87	-	10.7	19.5	-	Anonymus
macrophthalmus	Aegean and Mediterranean Sea													(1993)
Diplodus	Edremit Bay	F	1997-98	C	FL	0.0367	2.797	0.865*	322	10.33	7.3	13.8	0.57	Koc et al.
annularis														(2002)
Diplodus	Edremit Bay	M	1997-98	C	FL	0.0455	2.696	0.846*	330	10.18	8.0	14.0	0.55	Koc et al.
annularis														(2002)
Diplodus annularis	izmir Bay	C	1997-99	C	FL	0.014	3.190		160		7.8	15.6		Kinacigil & Akyol (2001)
Diplodus annularis	izmir Bay	F	1997-99	C	FL	0.018	3.090		72					Kinacigil & Akyol (2001)
Diplodus annularis	izmir Bay	M	1997-99	C	FL	0.012	3.260		68					Kinacigil & Akyol (2001)
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Diplodus annularis	dandarli Bay	C	1992	Wl	TL	0.04792	2.643	0.886*	24	-	-	-	-	Benli et al. (2000)
Diplodus annularis	Gulluk Bay	C	1992	Wl	TL	0.02056	3.016	0.957*	20	-	-	-	-	Benli et al. (2000)
Diplodus annularis	Gulluk Bay	C	1993	SP	TL	0.02791	2.926	0.945*	20	-	-	-	-	Benli et al. (2000)
Diplodus annularis	dandarli Bay	C	1993	AUT	TL	0.01509	3.187	0.927*	36	-	-	-	-	Benli et al. (2000)
Diplodus annularis	Izmir Bay	C	1993	AUT	TL	0.01449	3.196	0.931*	32	-	-	-	-	Benli et al. (2000)
Diplodus annularis	Gokova Bay	C	1993	AUT	TL	0.09692	2.473	0.893*	18	-	-	-	-	Benli et al. (2000)
Diplodus annularis	Gulbahoe Bay	C	19961997	C	FL	0.021	3.023	0.961	205		6.5	15.0		Tosunoglu et al. (1997)
Diplodus annularis	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.0000050	3.299	0.929*	214		8.7	17.3		Anonymus (1993)
Diplodus annularis	Marmara, Aegean and Mediterranean Sea	C	1991-93	SU	FL	0.0000248	2.959	0.895*	381		8.4	15.4		Anonymus (1993)
Diplodus annularis	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000327	2.905	0.918*	193		8.2	19.3		Anonymus (1993)
Diplodus annularis	Marmara, Aegean and Mediterranean Sea	C	1991-93	Wl	FL	0.0000102	3.150	0.976*	104		9.1	16.6		Anonymus (1993)
Diplodus sargus	Iskenderun Bay	C	2000	AUT	TL	0.0342	2.807	0.850	33	19.29	14.90	26.7	2.94	Can et al. (2000)
Diplodus vulgaris	Saros Bay	C	1992	Wl	TL	0.02840	2.929	0.943*	20	-	-	-	-	Benli et al. (2000)
Diplodus vulgaris	Izmir Bay	C	1993	AUT	TL	0.02413	2.983	0.952*	35	-	-	-	-	Benli et al. (2000)
Diplodus vulgaris	Iskenderun Bay	C	2000	AUT	TL	0.0131	3.124	0.933	105	16.66	13.20	27.0	2.36	Can et al. (2000)
Diplodus vulgaris	Marmara, Aegean and Mediterranean Sea	C	1991-93	Wl	FL	0.0000334	2.929	0.943*	20		12.1	17.5		Anonymus (1993)
Diplodus vulgaris	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.000365	2.908	0.992*	20		10.5	18.9		Anonymus (1993)
Diplodus vulgaris	Marmara, Aegean and Mediterranean Sea	C	1991-93	SU	FL	0.0000680	2.795	0.984*	11		8.8	16.7		Anonymus (1993)
Diplodus vulgaris	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000016	3.538	0.960*	18		13.0	18.0		Anonymus (1993)
Dipturus oxyrinnchus	Sigacik trawl area	C	2003	Wl, SU	TL	0.0007	3.40	0.99	8	-	17.9	62.2	-	Filiz & Bilge (2004)
Engraulis encrasicolus	Eastern Black Sea	C	2001-02	Wl	SL	0.032	2.507	0.895	50	10.48	9.45	11.7	0.066	Aka (2003)
Engraulis encrasicolus	Mid Black Sea	C	2001-02	Wl	SL	0.0214	2.673	0.913	50	10.04	8.75	11.35	0.088	Aka (2003)
Engraulis encrasicolus	Western Black Sea	C	2001-02	Wl	SL	0.0111	2.979	0.865	50	10.28	9.35	11.05	0.061	Aka (2003)
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Engraulis	Marmara Sea	C	2001-02	WI	SL	0.0094	2.999	0.942	50	11.34	10.5	12.1	0.060	Aka (2003)
encrasicolus														
Engraulis	Izmir Bay	C	2001-02	WI	SL	0.0102	2.937	0.852	50	10.34	9.5	11.2	0.049	Aka (2003)
encrasicolus														
Engraulis	Edremit Bay	C	2001-02	WI	SL	0.009	2.991	0.866	50	10.14	9.15	11.45	0.062	Aka (2003)
encrasicolus														
Engraulis	Eastern Black	F	1997-98	-	TL	0.0054	3.040	0.891	907	11.71	8.8	13.8	3.42	Gozler
encrasicolus	Sea													& Ciloglu (1998)
Engraulis	Eastern Black	M	1997-98	-	TL	0.0049	3.071	0.640	339	11.68	8.8	13.8	1.74	Gozler
encrasicolus	Sea													& Ciloglu (1998)
Engraulis	Eastern Black	C	1997-98	-	TL	0.0057	3.015	0.855	1664	11.22	7.0	13.8	4.43	Gozler
encrasicolus	Sea													& Ciloglu (1998)
Engraulis encrasicolus	Eastern Black Sea	C	1996-97	-	TL	0.00569	3.117	-	-	-	6.2	13.5	-	Kayali (1998)
Engraulis encrasicolus	Eastern Black sea	C	1993-94		TL	0.0051	3.048			10.43				Duzgunes et al. (1995)
Engraulis encrasicolus	Black Sea	C	1988-94	C	TL	0.0053	3.038	0.941*	43					Erkoyuncu et al. (1994)
Engraulis	Black Sea	C	-	-	-	0.0047	3.100	-	-	-	-	-	-	Ozdamar
encrasicolus														et al. (1991a)
Engraulis	Black Sea	C	-	-	-	0.023	3.412	-	-	-	-	-	-	Ozdamar
encrasicolus														et al. (1991b)
Engraulis	Black Sea	C	-	-	-	0.0025	3.388	-	-	-	-	-	-	Karacam
encrasicolus														& Duzgunes (1990)
Engraulis	Black Sea	C	1987-88	-	TL	-2.5941	3.386	-	956	9.34	6.66	15.25	-	Duzgunes
encrasicolus														& Karacam (1989)
Engraulis	Black Sea	C	1988-89	C	TL	0.00643	2.974	-	1172	-	75	130	-	Unsal
encrasicolus														(1989)**
Epinephelus	Iskenderun	C	2000	AUT	TL	0.0885	2.391	0.930	53	29.76	14.20	55.4	8.78	Can et al.
costae	Bay													(2000)
Gadus euxinus	Eastern Black Sea	C				0.0272	2.573							Duzgunes & Karacam (1990)
Gadus merlangus	Black Sea	C	1995-96	C	TL	0.0039	3.238	0.931	1302	14.53	9.0	24.0	0.07	Samsun &
euxinus														Erkoyuncu (1998)
Gadus merlangus	Black Sea	C	1988-94	C	TL	0.0034	3.299	0.922*	54	-	-	-	-	Erkoyuncu
euxinus														et al. (1994)
Gadus merlangus	Mid Black	C	-	C	-	00043	3.195	-	-	-	-	-	--	Samsun
euxinus	Sea													et al. (1993)
Gadiculus	Sigacik trawl	C	2003	C	TL	0.0056	3.24	0.89	110	-	6.4	10.5	-	Filiz
argenteus argenteus	area													& Bilge (2004)
Gymnura	Sigacik trawl	C	2003	C	TL	0.0268	2.96	0.98	9	-	37.5	72.0	-	Filiz
alatvela	area													& Bilge (2004)
Helicolenus	Sigacik trawl	C	2003	C	TL	0.0079	3.28	0.92	178	-	5.5	13.5	-	Filiz
dactylopterus	area													& Bilge (2004)
Hoplostethus mediterraneus	Sigacik trawl area	C	2003	C	TL	0.0149	2.95	0.98	137		8.0	18.0		Filiz & Bilge (2004)
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Lesueurigobius	Sigacik trawl	C	2003	C	TL	0.0392	2.13	0.72	17	-	6.2	8.1	-	Filiz
friessi	area													& Bilge (2004)
Lepidopus	Sigacik trawl	C	2003	C	TL	0.0004	3.11	0.99	40	-	21.9	81.5	-	Filiz
caudatus	area													& Bilge (2004)
Liza ramada	Gulluk Lagoon	C	1993-94	C	TL	0.0066894	3.052	0.946*	86	23.5	17.0	33.8	2.8	Hossucu (2001)
Liza saliens	Gulluk Lagoon	C	1993-94	C	TL	0.008859	2.997	0.965*	38	25.3	18.3	39.0	3.7	Hossucu (2001)
Liza saliens	Izmir Bay	U	1997-98	C	FL	0.0082	3.090	0.990	15	-	8.4	19.6	3.29	Akyol
														(1999)
Liza saliens	Izmir Bay	F	1997-98	C	FL	0.0088	3.080	0.953	294	-	13.4	33.1	-	Akyol
														(1999)
Liza saliens	Izmir Bay	M	1997-98	C	FL	0.0079	3.120	0.986	121	-	13.2	28.7	-	Akyol
														(1999)
Liza saliens	Izmir Bay	C	1997-98	C	FL	0.0078	3.120	0.985	430	-	8.4	33.1	-	Akyol
														(1999)
Macroramphosus	Sigacik trawl	C	2003	C	TL	0.0079	3.28	0.92	43	-	7.1	11.4	-	Filiz
scolopax	area													& Bilge (2004)
Merlangius	Black Sea	F	1990-93	C	-	0.0040	3.250	0.980*	4176	-	-	-	-	l§men
merlangus														(2002)
euxinus														
Merlangius	Black Sea	M	1990-93	C	-	0.0044	3.220	0.980*	3181	-	-	-	-	l§men
merlangus														(2002)
euxinus														
Merlangius	Eastern Black	F	1997	C	-	0.004856	3.151	-	1349	-	-	-	-	Sahin
merlangus	Sea													& Akbulut
euxinus														(1997)
Merlangius	Eastern Black	M	1997	C	-	0.005450	3.110	-	864	-	-	-	-	Sahin
merlangus	Sea													& Akbulut
euxinus														(1997)
Merlangius	Eastern Black	C	1990	C	-	0.0070	3.007	-	-	-	-	-	-	Uysal
merlangus	Sea													(1990)
euxinus														
Merluccius	Saros Bay	C	1991	SU	TL	0.00669	3.013	0.736*	275	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	dandarli Bay	C	1991	SU	TL	0.00933	2.617	0.64*	78	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	Izmir Bay	C	1991	SU	TL	0.01038	2.862	0.674*	53	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	Gokova Bay	C	1991	SU	TL	0.00732	2.977	0.681*	203	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	Saros Bay	C	1992	WI	TL	0.00382	3.221	0.960*	74	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	dandarli Bay	C	1992	WI	TL	0.00507	3.097	0.986*	156	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	Izmir Bay	C	1992	WI	TL	0.00383	3.188	0.995*	35	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	Gokova Bay	C	1992	WI	TL	0.00573	3.025	0.997*	21	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	Saros Bay	C	1992	SP	TL	0.00297	3.164	0.960*	170	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	dandarli Bay	C	1992	SP	TL	0.00572	3.029	0.941*	73	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	Izmir Bay	C	1992	SP	TL	0.00670	3.101	0.980*	9	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	Gokova Bay	C	1992	SP	TL	0.00570	3.050	0.960*	68	-	-	-	-	Benli et al.
merluccius														(2000)
Merluccius	Saros Bay	C	1993	SP	TL	0.00297	3.075	0.970*	60	-	-	-	-	Benli et al.
merluccius														(2000)
198
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Merluccius merluccius	dandarli Bay	C	1993	SP	TL	0.00561	3.069	0.991*	35	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Izmir Bay	C	1993	SP	TL	0.00665	3.037	0.903*	37	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Gulluk Bay	C	1993	SP	TL	0.00291	3.257	0.943*	17	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Gokova Bay	C	1993	SP	TL	0.00618	3.049	0.983*	51	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Saros Bay	C	1993	AUT	TL	0.00642	3.028	0.997*	76	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	dandarli Bay	C	1993	AUT	TL	0.00312	3.266	0.989*	68	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Izmir Bay	C	1993	AUT	TL	0.00432	3.175	0.986*	58	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Gulluk Bay	C	1993	AUT	TL	0.00731	3.017	0.993*	20	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Gokova Bay	C	1993	AUT	TL	0.00396	3.173	0.977*	26	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Saros Bay	C	1991	SU	TL	0.00669	3.013	0.736*	275	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	dandarli Bay	C	1991	SU	TL	0.00933	2.617	0.640*	78	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Izmir Bay	C	1991	SU	TL	0.01038	2.862	0.674*	53	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Gokova Bay	C	1991	SU	TL	0.00732	2.977	0.681*	203	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Saros Bay	C	1992	WI	TL	0.00382	3.221	0.960*	74	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	dandarli Bay	C	1992	WI	TL	0.00507	3.097	0.986*	156	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Izmir Bay	C	1992	WI	TL	0.00383	3.188	0.995*	35	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Gokova Bay	C	1992	WI	TL	0.00573	3.025	0.997*	21	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Saros Bay	C	1992	SP	TL	0.00297	3.167	0.960*	170	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	dandarli Bay	C	1992	SP	TL	0.00572	3.029	0.941*	73	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Izmir Bay	C	1992	SP	TL	0.00670	3.014	0.980*	9	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Gokova Bay	C	1992	SP	TL	0.00570	3.050	0.960*	68	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Saros Bay	C	1993	SP	TL	0.00572	3.075	0.970*	60	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	dandarli Bay	C	1993	SP	TL	0.00561	3.069	0.991*	35	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Izmir Bay	C	1993	SP	TL	0.00665	3.037	0.903*	37	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Gulluk Bay	C	1993	SP	TL	0.00291	3.257	0.943*	17	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Gokova Bay	C	1993	SP	TL	0.00618	3.049	0.983*	51	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Saros Bay	C	1993	AUT	TL	0.00642	3.028	0.977*	76	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	dandarli Bay	C	1993	AUT	TL	0.00312	3.266	0.989*	68	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Izmir Bay	C	1993	AUT	TL	0.00432	3.175	0.983*	58	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Gulluk Bay	C	1993	AUT	TL	0.00731	3.017	0.993*	20	-	-	-	-	Benli et al. (2000)
Merluccius merluccius	Gokova Bay	C	1993	AUT	TL	0.00396	3.173	0.977*	26	-	-	-	-	Benli et al. (2000)
200
ANNALES • Ser. hist. nat. • 14 • 2004 • 2
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Merluccius merluccius	Aegean Sea	C	1994-95	C	TL	0.0045	3.194	0.974*	336		13.6	43.5		Uckun et al. (2000)
Merluccius merluccius	Edremit Bay	C	1996-97		TL	0.0095	2.928	0.960*	165		15.8	37.2		Torcu et al. (1997)
Merluccius merluccius	Marmara, Aegean and Mediterranean Sea	C	1991-93	SU	FL	0.0000048	3.065	0.984*	1164		10.5	69.5		Anonymus (1993)
Merluccius merluccius	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000037	3.117	0.990*	1114		7.8	73.5		Anonymus (1993)
Merluccius merluccius	Marmara, Aegean and Mediterranean Sea	C	1991-93	WI	FL	0.0000043	3.090	0.988*	614		14.2	79.0		Anonymus (1993)
Merluccius merluccius	Izmir Bay	C	1989	AUT	FL	0.00246	2.960		74	26.10	20.50	36.14		Kara & Gurbet (1990)
Mugil cephalus	Gulluk Lagoon	C	1993-94	C	TL	0.008198	3.048	0.976*	132	31.0	17.6	46.2	6.6	Hossucu (2001)
Mugil cephalus	Izmir Bay	U	1997-98	C	FL	0.0149	2.950	0.998	29	-	10.2	45.3	-	Akyol (1999)
Mugil cephalus	Izmir Bay	F	1997-98	C	FL	0.0075	3.160	0.989	98	-	25.7	54.5	-	Akyol (1999)
Mugil cephalus	Izmir Bay	M	1997-98	C	FL	0.0109	3.040	0.965	77	-	28.2	55.5	-	Akyol (1999)
Mugil cephalus	Izmir Bay	C	1997-98	C	FL	0.0101	3.070	0.991	204	-	10.2	55.5	-	Akyol (1999)
Mugil so-iuy	Eastern Black Sea	C	1995	C	TL	0.010	2.980	0.970*	174		22.5	68.9		Basoinar & Okumus (1997)
Mugil sp.	Black Sea	C	1988-94	C	TL	0.0068	3.077	0.980*	75					Erkoyuncu et al. (1994)
Mullus barbatus	Izmir Bay	C	1997	WI	FL	0.0071	3.290		221		10.3	19.1		Kinacigil et al. (2001)
Mullus barbatus	Saroz Bay	C	1991	SU	TL	0.01493	2.968	0.965*	222	-	-	-	-	Benli et al. (2000)
Mullus barbatus	dandarli Bay	C	1991	SU	TL	0.00775	3.234	0.918*	40	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Izmir Bay	C	1991	SU	TL	0.01540	2.988	0.905*	118	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Gulluk Bay	C	1991	SU	TL	0.02774	2.768	0.704*	122	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Gokova Bay	C	1991	SU	TL	0.01578	2.998	0.857*	60	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Saroz Bay	C	1992	WI	TL	0.00942	3.188	0.977*	80	-	-	-	-	Benli et al. (2000)
Mullus barbatus	dandarli Bay	C	1992	WI	TL	0.001261	3.067	0.973*	166	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Izmir Bay	C	1992	WI	TL	0.00871	3.204	0.980*	39	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Gulluk Bay	C	1992	WI	TL	0.01442	3.051	0.980*	53	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Gokova Bay	C	1992	WI	TL	0.00824	3.219	0.967*	40	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Saroz Bay	C	1992	SP	TL	0.00321	3.309	0.922*	170	-	-	-	-	Benli et al. (2000)
200
ANNALES • Ser. hist. nat. • 14 • 2004 • 2
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Mullus barbatus	dandarli Bay	C	1992	SP	TL	0.00328	3.313	0.960*	100	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Izmir Bay	C	1992	SP	TL	0.00346	3.305	0.980*	120	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Gulluk Bay	C	1992	SP	TL	0.00110	3.051	0.941*	60	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Gokova Bay	C	1992	SP	TL	0.04830	2.801	0.884*	11	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Saroz Bay	C	1993	SP	TL	0.01421	3.029	0.990*	38	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Izmir Bay	C	1993	SP	TL	0.01382	3.038	0.938*	91	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Gulluk Bay	C	1993	SP	TL	0.01205	3.115	0.981*	40	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Gokova Bay	C	1993	SP	TL	0.00736	3.314	0.960*	68	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Saroz Bay	C	1993	AUT	TL	0.03087	2.738	0.899*	48	-	-	-	-	Benli et al. (2000)
Mullus barbatus	dandarli Bay	C	1993	AUT	TL	0.02738	2.824	0.963*	40	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Izmir Bay	C	1993	AUT	TL	0.01723	2.984	0.902*	115	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Gokova Bay	C	1993	AUT	TL	0.00767	3.292	0.950*	99	-	-	-	-	Benli et al. (2000)
Mullus barbatus	Iskenderun Bay	C	1992-93	C	FL	-1.77494	3.026	0.956*	348		7.5	15.12		Türeli & Erdem (1997)
Mullus barbatus	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.0000033	3.320	0.960*	533		8.5	22.4		Anonymus (1993)
Mullus barbatus	Marmara, Aegean and Mediterranean Sea	C	1991-93	Wl	FL	0.0000091	3.106	0.970*	636		9.0	20.5		Anonymus (1993)
Mullus barbatus	Marmara, Aegean and Mediterranean Sea	C	1991-93	SU	FL	0.00000201	2.941	0.955*	1084		8.5	21.0		Anonymus (1993)
Mullus barbatus	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000078	3.143	0.968*	1090		8.8	22.3		Anonymus (1993)
Mullus barbatus	Mid Black sea	C	-	-	-	0.0054	3.209	-	-	-	-	-	-	Anonymus (1991a)
Mullus barbatus	Izmir Bay	C	1989	AUT	FL	0.00542	3.344		231	13.30	10.83	19.50		Kara & Gurbet (1990)
Mullus barbatus	Aliaga-dandarli Bay	C	1989	AUT	FL	0.13243	2.310		199	14.68	11.20	19.12		Kara & Gurbet (1990)
Mullus barbatus	Edremit Bay	C	1989	AUT	FL	0.011782	2.960		196	15.34	11.50	18.75		Kara & Gurbet (1990)
Mullus barbatus	Gulbahoe Bay (Izmir)	C	1973	C	FL	0.0165	2.923	0.960*	6054	-	7.6	22.0	-	T°gulga (1977)
Mullus barbatus	Edremit Bay	C	1970	SU	FL	0.022	-	-	204	-	10.0	19.0	-	Kinikaslan (1972)
Mullus barbatus ponticus	Eastern Black Sea	F	1997	C		0.0052653	3.217							Sahin & Akbulut (1997)
200
ANNALES • Ser. hist. nat. • 14 • 2004 • 2
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Mullus barbatus ponticus	Eastern Black Sea	M	1997	C		0.0053736	3.220							Sahin & Akbulut (1997)
Mullus barbatus ponticus	Black Sea	C	1988-89	C	TL	0.006855	3.156		2116	12.0	6.9	25.3	0.004	Samsun & Erkoyuncu (1992)
Mullus surmuletus	Saros Bay	C	1991	su	TL	0.0078	3.089	0.950*	57	-	-	-	-	Benli et al. (2000)
Mullus surmuletus	Gokova Bay	C	1991	su	TL	0.02168	2.920	0.926*	20	-	-	-	-	Benli et al. (2000)
Mullus surmuletus	Saros Bay	C	1992	Wl	TL	0.03224	2.819	0.830*	6	-	-	-	-	Benli et al. (2000)
Mullus surmuletus	Gulluk Bay	C	1992	Wl	TL	0.00984	3.242	0.913*	20	-	-	-	-	Benli et al. (2000)
Mullus surmuletus	dandarli Bay	C	1992	SP	TL	0.00510	3.221	0.810*	27	-	-	-	-	Benli et al. (2000)
Mullus surmuletus	Izmir Bay	C	1992	SP	TL	0.00580	3.194	0.980	28	-	-	-	-	Benli et al. (2000)
Mullus surmuletus	Izmir Bay	C	1993	AUT	TL	0.01177	3.131	0.958	37	-	-	-	-	Benli et al. (2000)
Mullus surmuletus	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.0000087	3.122	0.947*	170		11.5	32.1		Anonymus (1993)
Mullus surmuletus	Marmara, Aegean and Mediterranean Sea	C	1991-93	su	FL	0.0000488	2.784	0.929*	131		114.0	205.0		Anonymus (1993)
Mullus surmuletus	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000103	3.104	0.990*	67		97.0	238.0		Anonymus (1993)
Mullus surmuletus	Marmara, Aegean and Mediterranean Sea	C	1991-93	Wl	FL	0.0000209	2.971	0.968*	30		100.0	253.0		Anonymus (1993)
Mustelus mustelus	Sigacik trawl area	C	2003	Wl, SU	TL	0.0011	3.25	0.99	35		38.3	97.5		Filiz & Bilge (2004)
Myliobatis aquila	Sigacik trawl area	C	2003	Wl, SU	TL	0.0008	3.34	0.93	14		47.5	76.5		Filiz & Bilge (2004)
Oblada melanura	Iskenderun Bay	C	2000	AUT	TL	0.0322	2.697	0.964	22	20.45	15.60	27.0	3.32	Can et al. (2000)
Pagellus sp.	Izmir Bay	C	1989	AUT	FL	0.01592	3.008		111	14.59	8.66	21.43		Kara & Gurbet (1990)
Pagellus sp.	Edremit Bay	C	1989	AUT	FL	0.14723	2.289		74	16.75	13.71	22.0		Kara & Gurbet (1990)
Pagellus acerna	Aegean Sea	F	1991-93	C	FL	0.00527	3.447	0.850*	85	14.74	14.7	21.0	0.34	Ozaydin (1997)
Pagellus acerna	Aegean Sea	M	1991-93	C	FL	0.01087	3.172	0.927*	139	13.68	10.5	17.8	0.22	Ozaydin (1997)
Pagellus acerna	Aegean Sea	C	1991-93	C	FL	0.01420	3.075	0.962*	302	13.27	7.8	21.0	0.24	Ozaydin (1997)
Pagellus acerna	Gulbahoe Bay	C	1996-97	C	FL	0.082	3.288	0.935	107		9.8	14.8		Tosunoglu et al. (1997)
Pagellus acerna	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000876	2.646	0.806*	40		8.0	15.0		Anonymus (1993)
200
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Pagellus bogaraveo	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.0000018	3.472	0.951*	73		9.0	20.0		Anonymus (1993)
Pagellus bogaraveo	Marmara, Aegean and	C	1991-93	SU	FL	0.0000083	3.150	0.978*	5	-	11.0	15.0	-	Anonymus (1993)
Pagellus bogaraveo	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000709	2.709	0.792*	20		11.0	14.0		Anonymus (1993)
Pagellus bogaraveo	Marmara, Aegean and Mediterranean Sea	C	1991-93	WI	FL	0.0000061	3.214	0.968*	81		9.0	19.0		Anonymus (1993)
Pagrus coeruleostictus	iskenderun Bay	C	2000	AUT	TL	0.0671	2.521	0.908	311	18.52	12.50	38.8	2.93	Can et al. (2000)
Pagellus erythinus	Saros Bay	C	1991	SU	TL	0.01299	2.972	0.832*	124	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	dandarli Bay	C	1991	SU	TL	0.01190	3.156	0.959*	20	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	izmir Bay	C	1991	SU	TL	0.04097	2.889	0.857*	60	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Gulluk Bay	C	1991	SU	TL	0.01816	2.968	0.908*	40	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Gokova Bay	C	1991	SU	TL	0.0255	2.932	0.822*	40	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Saros Bay	C	1992	WI	TL	0.03184	2.773	0.896*	20	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	dandarli	C	1992	WI	TL	0.01459	3.079	0.978*	29	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Gokova Bay	C	1992	WI	TL	0.01356	3.101	0.945*	20	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Saros Bay	C	1992	SP	TL	0.00109	3.537	0.941*	46	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	dandarli Bay	C	1992	SP	TL	0.01619	3.011	0.960*	40	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	izmir Bay	C	1992	SP	TL	0.06402	2.752	0.960*	56	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Gulluk Bay	C	1992	SP	TL	0.02065	2.527	0.689*	42	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Gokova Bay	C	1992	SP	TL	0.04277	2.838	0.960*	7	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Saros Bay	C	1993	SP	TL	0.01249	3.140	0.988*	16	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	izmir Bay	C	1993	SP	TL	0.01498	3.077	0.988*	57	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Gulluk Bay	C	1993	SP	TL	0.06999	2.539	0.975*	17	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Gokova Bay	C	1993	SP	TL	0.01754	2.993	0.956*	17	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Saros Bay	C	1993	AUT	TL	0.00606	3.391	0.984*	17	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	dandarli Bay	C	1993	AUT	TL	0.06999	2.539	0.975*	17	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	izmir Bay	C	1993	AUT	TL	0.01699	3.001	0.944*	57	-	-	-	-	Benli et al. (2000)
Pagellus erythinus	Aegean Sea	F	1991-93	C	FL	0.02370	2.907	0.978*	512	15.36	11.1	24.5	0.18	Ozaydin (1997)
Pagellus erythinus	Aegean Sea	M	1991-93	C	FL	0.2722	2.865	0.956*	163	18.27	8.8	25.6	0.41	Ozaydin (1997)
Pagellus erythinus	Aegean Sea	C	1991-93	C	FL	0.02423	2.985	0.972*	692	16.04	8.8	25.6	0.20	Ozaydin (1997)
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Pagellus erythinus	Gulbahoe Bay	C	1996-97	C	FL	0.018	3.012	0.976	204		8.0	24.0		Tosunoglu et al. (1997)
Pagellus erythinus	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000287	2.917	0.966*	387		7.0	25.0		Anonymus (1993)
Pagellus erythinus	Marmara, Aegean and Mediterranean Sea	C	1991-93	Wl	FL	0.0000122	3.080	0.966*	117		11.0	24.0		Anonymus (1993)
Peristedion cataphractum	Sigacil trawl area	C	2003	C	TL	0.0048	2.97	0.99	11		8.1	21.2		Filiz & Bilge (2004)
Pleuronactes platessa	Black Sea	C	1988-94	C	TL	0.0078	3.109	0.960*	48					Erkoyuncu et al. (1994)
Phycis blennoides	Sigacik trawl area	C	2003	C	TL	0.0017	3.55	0.89	12		12.3	15.0		Filiz & Bilge (2004)
Pomadasys incisus	iskenderun Bay	C	2000	AUT	TL	0.0465	2.604	0.910	106	16.12	13.40	21.2	1.75	Can et al. (2000)
Pomatomus saltatrix	Black Sea	C	1988-94	C	TL	0.0388	2.558	0.846*	19					Erkoyuncu et al. (1994)
Pomatomus saltator	izmir Bay	C	1987	AUT -Wl	TL	0.02017	2.955		400		15.0	40.5		Alpbaz & Kinacigil (1988)
Raja clavata	Sinop	C	-	-	-	0.0026	3.200	0.990	54	-	27.3	83.2	-	Erdem et al. (2001)
Raja clavata	Sigacik trawl area	C	2003	Wl, SU	TL	0.0016	3.30	0.94	37		20.5	99.0		Filiz & Bilge (2004)
Raja miraletus	Sigacik trawl area	C	2003	Wl, SU	TL	0.0001	4.15	0.93	13		30.0	50.5		Filiz & Bilge (2004)
Raja sp.	Black Sea	C	1988-94	C	TL	0.0090	2.920	0.922*	40					Erkoyuncu et al. (1994)
Sarda sarda	Black Sea	C	1995-96	AUT -Wl	TL	0.0058	3.176	0.980*	4104	34.42	20.0	40.0	2.21	Samsun (1997)
Sarda sarda	Black Sea	C	1988-94	C	TL	0.0297	2.679	0.865*	14					Erkoyuncu et al. (1994)
Sardina pilchardus	izmir Bay	F	1996-97	C	TL	0.0053	3.275	0.872*	187		9.3	13.0		Karakayis & Togulga (2000)
Sardina pilchardus	izmir Bay	M	1996-97	C	TL	0.0081	3.108	0.781*	75					Karakayis & Togulga (2000)
Sardina pilchardus	izmir Bay	C	1996-97	C	TL	0.0062	3.214	0.855*	262		9.3	14.6		Karakayis & Togulga (2000)
Sardina pilchardus	izmir Bay	F	1997-98	C	FL	0.0042	3.382	0.850	216		9.6	14.7		Mater & Bayhan (1999)
Sardina pilchardus	izmir Bay	M	1997-98	C	FL	0.0025	3.600	0.870	148		9.8	14.9		Mater & Bayhan (1999)
Sardina pilchardus	izmir Bay	C	1997-98	C	FL	0.0045	3.359	0.830	364		9.6	14.9		Mater & Bayhan (1999)
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Saurida undosquamis	iskenderun	F	1999-00	C	TL	0.088	3.190	0.980	368	-	9.0	35.0	-	Ismen (2002)
Saurida undosquamis	iskenderun	M	1999-00	C	TL	0.120	2.950	0.960	234	-	5.0	32.0	-	Ismen (2002)
Saurida undosquamis	Fethiye Bay	C	1993	C	FL	-1.3279	3.295	0.828*	430		17.3	30.8		Mater & Torcu (1997)
Saurida undosquamis	Mersin Bay	C	1993	C	FL	-0.9596	2.616	0.656*	100		13.5	32.2		Mater & Torcu (1997)
Saurida undosquamis	iskenderun Bay	C	1992-93	C	FL	-2.06171	3.022	0.998*	333		8.3	21.92		Türeli & Erdem (1997)
Scomberomorus commerson	Gulluk and Gokova Bay	C	1994	C	TL	0.1567	2.223	0.885*	70		52.0	87.0		Buhan et al. (1997)
Scophthalmus maeaticus	Black Sea	C	-	C	-	0.0047	3.418	-	-	-	40.0	-	-	Erdem (2001)
Scophthalmus maximus	Eastern Black Sea	F	1990-91	C	TL	0.0099	3.140	0.980	86	-	196	630	-	Avsar (1999)**
Scophthalmus maximus	Eastern Black Sea	M	1990-91	C	TL	0.0112	3.110	0.980	63	-	181	595	-	Avsar (1999)**
Scophthalmus maximus	Eastern Black Sea	C	1990-91	C	TL	0.0085	3.180	0.980	194	-	181	630	-	Avsar (1999)**
Scorpaena porcus	Sinop	C	1996-97	-	TL	0.054	2.590	-	633	17.26	11.49	23.63	0.083	Koca (2002)
Scorpaena sp.	Black Sea	C	1988-94	C	TL	0.0180	3.080	0.980*	31					Erkoyuncu et al. (1994)
Scyliorhinus canicula	Northern Aegean Sea	F	1991-96	SP	TL	0.001	3.453	0.991	95			51.7		Cihangir et al. (1997)
Scyliorhinus canicula	Northern Aegean Sea	M	1991-96	SP	TL	0.005	2.004	0.989	121			54.6		Cihangir et al. (1997)
Scyliorhinus canicula	Northern Aegean Sea	C	1991-96	SU-AUT	TL	0.001	3.205	0.919	370			54.6		Cihangir et al. (1997)
Scyliorhinus canicula	Siagacik trwal area	C	2003	Wl, SU	TL	0.0012	3.26	0.99	637	-	10.5	50.9	-	Filiz & Bilge (2004)
Serranus cabrilla	Edremit Bay	C	1997-98	C	FL	0.0311	2.670	0.880	595	14.43	8.6	22.3	1.573	Torcu et al. (2004
Serranus cabrilla	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.0000225	2.879	0.937*	100		94.0	200.0		Anonymus (1993)
Serranus cabrilla	Marmara, Aegean and Mediterranean Sea	C	1991-93	SU	FL	0.0000081	3.059	0.929*	193		94.0	255.0		Anonymus (1993)
Serranus cabrilla	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000066	3.121	0.966*	176		10.4	20.3		Anonymus (1993)
Serranus cabrilla	Marmara, Aegean and Mediterranean Sea	C	1991-93	Wl	FL	0.00001	3.043	0.980*	75		11.6	17.5		Anonymus (1993)
Serranus scriba	Marmara, Aegean and Mediterranean Sea	C	1991-93	SU	FL	0.0000252	2.878	0.719	20		8.0	24.0		Anonymus (1993)
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Siganus rivulatus	Antalya Bay	F	1996-98	C	TL	0.0064	3.221	0.903*	292	15.12	7.0	21.5	0.32	Bilecenoglu & Kaya (2002)
Siganus rivulatus	Antalya Bay	M	1996-98	C	TL	0.007945	3.135	0.903*	229	16.09	7.1	20.6	0.34	Bilecenoglu & Kaya (2002)
Siganus rivulatus	Antalya Bay	C	1996-98	C	TL	0.007137	3.179	0.903*	521		7.0	21.5		Bilecenoglu & Kaya (2002)
Solea solea	iskenderun Bay	F	2000-01	C	TL	0.0093	3.077	0.960	530	-	20.5	28.2	-	Türkmen (2003)
Solea solea	iskenderun Bay	M	2000-01	C	TL	0.0117	2.998	0.973	553	-	8.8	25.0	-	Türkmen (2003)
Solea sp.	Black Sea	C	1988-94	C	TL	0.0019	3.580	0.941*	19					Erkoyuncu et al. (1994)
Solea solea	izmir Bay	C	1989-90	C	TL	0.00514	3.135	0.815	335	26.2	11.0	34.5	0.6	Hossucu (1992)
Solea solea	izmir Bay	C	1989	AUT	FL	0.00054	3.640		9	23.61	22.50	25.83		Kara & Gurbet (1990)
Solea solea	AliaUa-dan-darli Bay	C	1989	AUT	FL	0.00182	3.458		11	25.31	20.50	30.10		Kara & Gurbet (1990)
Solea solea	Edremit Bay	C	1989	AUT	FL	0.20137	1.951		14	26.21	21.50	31.00		Kara & Gurbet (1990)
Solea vulgaris	Yumurtalik	C	1992-93	C	TL	-2.52566	3.250	0.992*	603		12.1	34.5		Altun et al. (1997)
Solea vulgaris	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.0000003	3.562	0.951*	8		23.8	34.2		Anonymus (1993)
Solea vulgaris	Marmara, Aegean and Mediterranean Sea	C	1991-93	SU	FL	0.0000026	3.183	0.937*	66		13.9	27.4		Anonymus (1993)
Solea vulgaris	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0001367	2.479	0.740*	23		19.2	33.5		Anonymus (1993)
Solea vulgaris	Marmara, Aegean and Mediterranean Sea	C	1991-93	Wl	FL	0.0000098	2.958	0.960*	11					Anonymus (1993)
Sparus aurata	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.000025	2.941	0.960*	24		15.5	24.1		Anonymus (1993)
Sparus aurata	Marmara, Aegean and Mediterranean Sea	C	1991-93	SU	FL	0.0000693	2.743	0.984*	9		15.3	22.2		Anonymus (1993)
Sparus aurata	Marmara, Aegean and Mediterranean Sea	C	1991-93	Wl	FL	0.000017	3.470	0.982*	31		14.2	22.3		Anonymus (1993)
Sparus aurata	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000149	3.023	0.878*	14		16.0	19.6		Anonymus (1993)
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Sphyraena chrysotaenia	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000053	3.049	0.895*	6		21.0	25.2		Anonymus (1993)
Sphyraena chrysotaenia	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.000049	3.056	0.964*	22		14.0	25.6		Anonymus (1993)
Sphyraena sphyraena	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.0001309	2.428	0.925*	9		24.0	34.0		Anonymus (1993)
Sphyraena sphyraena	Marmara, Aegean and Mediterranean Sea	C	1991-93	SU	FL	0.0002720	2.290	0.927*	13		22.0	35.0		Anonymus (1993)
Spicara flexuosa	Izmir Bay	F	1998-99	C	FL	0.1156	2.163	0.689*	240		9.20	14.90		Mater et al. (2001)
Spicara flexuosa	Izmir Bay	M	1998-99	C	FL	0.0452	2.566	0.740*	172		11.30	15.50		Mater et al. (2001)
Spicara flexuosa	Izmir Bay	C	1998-99	C	FL	0.0411	2.594	0.846*	412		9.20	15.50		Mater et al. (2001)
Spicara smaris	Eastern Black Sea	F	1989	C	TL	0.0510	3.229	0.958*	280	-	11.1	22.5	-	Sahin & Gene (1999)
Spicara smaris	Eastern Black Sea	M	1989	C	TL	0.0650	3.126	0.880*	176	-	11.3	22.0	-	Sahin & Gene (1999)
Spicara smaris	Eastern Black Sea	C	1991-92	C	TL	0.005	3.260	-	-	-	-	-	-	Ismen (1995)
Spicara smaris	Black Sea	C	1988-94	C	TL	0.0061	3.215	0.941*	25					Erkoyunueu et al. (1994)
Spicara smaris	Mid Black Sea	C	-	-	-	0.0051	3.217	-	-	-	-	-	-	Anonymus (1991e)
Sprattus sprattus phallericus	Eastern Black Sea	F	1991	SP-SU	TL	0.00214	3.456	0.999*	214	-	8.45	14.09		Sahin (1999)
Sprattus sprattus phallericus	Eastern Black Sea	M	1991	SP-SU	TL	0.00204	3.474	0.994*	158	-	8.27	12.5	-	Sahin (1999)
Sprattus sprattus phallericus	Eastern Black Sea	C	1991	SP-SU	TL	0.00211	3.461	0.997*	372	-	8.35	13.20	-	Sahin (1999)
Squalus acanthias	Black Sea	F	1969-73		TL	0.0041	3.004	0.996	1840	85	30.0	140.0		Kutaygil & Bileeik (1998)
Squalus acanthias	Black Sea	M	1969-73	C	TL	0.0053	2.929	0.999	1780	75	30.0	120.0		Kutaygil & Bileeik (1998)
Squalus acanthias	Eastern Black Sea	C	1994-95	C	TL	0.0013	3.254	0.81*	159		35.5	141.5		Karaeam et al. (1996)
Squalus acanthias	Sigacik trawl area	C	2003	Wl, SU	TL	0.0031	3.11	0.98	32		27.0	70.5		Filiz & Bigle (2004)
Symphodus tinca	Iskenderun Bay	C	2000	AUT	TL	0.0021	3.675	0.997	10	14.41	12.10	17.2	1.75	Can et al. (2000)
Thunnus thynnus	Aegean and Mediterranean Sea	C	1996-98	C	TL	0.0000181	2.988	0.994*	3577	118.40	59.0	275.0	37.0	Karakulak & Oray (2001)
Thunnus thynnus	Aegean and Mediterranean Sea	C	1992-94	C	FL <120	-21.780	0.409	0.931	145					Karakulak & Oray (1994)
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Thunnus thynnus	Aegean and Mediterranean Sea	C	1992-94	C	FL >120	-1 86.521	1.633	0.780	1494					Karakulak & Oray (1994)
Torpedo marmorata	Siagacik trawl area	C	2003	Wl, su	TL	0.0273	2.91	0.98	27		9.2	34.0		Filiz & Bilge (2004)
Trachurus mediterraneus	Eastern Black Sea	C	1996-97	-	TL	0.0108	2.980	-	-	-	6.3	17.8	-	Kayali (1998)
Trachurus mediterraneus	Mid Black sea	C	-	-	-	0.0089	2.950	-	-	-	-	-	-	Anonymus (1991b)
Trachurus mediterraneus	Black Sea	C				0.3612	1.160							Düzgünes & Karaoam (1991)
Trachurus mediterraneus ponticus	Black Sea	C			FL	0.004834	3.218		601		6.5	17.5		Sahin et al. (1997)
Trachurus trachurus	Mid Black Sea	C	1995-96	C	TL	0.00759	3.05		720	14.13	9.4	16.8	0.07	Yücel & Erkoyuncu (2000)
Trachurus trachurus	Black Sea	C	1988-94	C	TL	0.0290	2.485	0.960	77					Erkoyuncu et al. (1994)
Trachurus trachurus	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.0000288	3.832	0.976*	259		9.0	29.6		Anonymus (1993)
Trachurus trachurus	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000071	3.072	0.955*	496		8.1	38.5		Anonymus (1993)
Trachurus trachurus	Marmara, Aegean and Mediterranean Sea	C	1991-93	Wl	FL	0.0000139	3.964	0.974*	311		10.7	27.1		Anonymus (1993)
Trigla lucerna	Yumurtalik	C	1992-93	-	TL	-2.18077	3.088	0.966*	348	-	14.4	26.9	-	Altun et al. (1997)
Trigla lucerna	Black Sea	C				0.0085	3.032							Erdem et al. (1994)
Trigla lucerna	Black Sea	C	1988-94	C	TL	0.0070	3.089	0.980*	55					Erkoyuncu et al. (1994)
Trisopterus minutus	dandarli Bay	C	1993	SP	TL	0.0979	3.017	0.951*	35	-	-	-	-	Benli et al. (2000)
Trisopterus minutus	dandarli Bay	C	1993	AUT	TL	0.00709	3.144	0.910*	117	-	-	-	-	Benli et al. (2000)
Upeneus moluccensis	Gokova Bay	C	1993	AUT	TL	0.00606	3.351	0.962*	39	-	-	-	-	Benli et al. (2000)
Upeneus moluccensis	Mediterranean Sea	F	1991-92	C	FL	0.01051	3.150	0.945*	535	13.69	8.6	17.8	1.635	Kaya et al. (1999)
Upeneus moluccensis	Mediterranean Sea	M	1991-92	C	FL	0.00607	3.352	0.922*	176	11.16	8.5	16.1	1.378	Kaya et al. (1999)
Upeneus moluccensis	Fethiye Bay	C	1990-93		FL	-1.2889	3.386	0.828*	536		9.0	17.0		Torcu & Mater (1997)
Upeneus moluccensis	Mersin Bay	C	1990-93		FL	-1.0554	2.903	0.723*	500		8.9	16.0		Torcu & Mater (1997)
Upeneus moluccensis	Marmara, Aegean and Mediterranean Sea	C	1991-93	SP	FL	0.0000043	3.257	0.976*	152		81.0	167.0		Anonymus (1993)
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Species	Area	Sex	Year	S	L	a	b	r2	N	mean	min	max	SD	Source
Upeneus moluccensis	Marmara, Aegean and Mediterranean Sea	C	1991-93	su	FL	0.0000036	3.300	0.947*	300		93.0	176.0		Anonymus (1993)
Upeneus moluccensis	Marmara, Aegean and Mediterranean Sea	C	1991-93	AUT	FL	0.0000257	2.896	0.956*	207		86.0	178.0		Anonymus (1993)
Upeneus moluccensis	Marmara, Aegean and Mediterranean Sea	C	1991-93	Wl	FL	0.0000027	3.353	0.941*	120		93.0	173.0		Anonymus (1993)
Upeneus pori	Babadilli-mani bight	F	1999-00	C	TL	0.0073	3.120	0.963	461		6.5	15.5		Cicek et al. (2002)
Upeneus pori	Babadilli-mani bight	M	1999-00	C	TL	0.0103	2.976	0.949	534		6.3	14.7		Cicek et al. (2002)
Zosterisessor ophiocephalus	Izmir Bay	C	1999-00	C	TL	0.0086	3.060	0.950	1066	-	8.0	23.3	-	Akyol (2003)
* r converted into r2
** length-weight relationship corresponding to mm, g
RAZMERJA MED DOLŽINO IN TEŽO RIB, ŽIVEČIH V TURŠKIH MORJIH: PREGLED
Hatice TORCU KOd & Zeliha AKA ERDOGAN University of Balikesir, Faculty of Science and Arts, Department of Biology, TR-10100 Balikesir, Turkey
Jakov DULČIČ
Inštitut za oceanografijo in ribištvo, HR-21000 Split, P.O. BOX 500, Hrvaška E-mail: dulcic@izor.hr
POVZETEK
Članek navaja 360 razmerij med dolžino in težo rib, in sicer glede na podatke, ki so bili zbrani iz obstoječe literature in zadevajo 90 vrst iz 40 družin, živečih v turških morjih. Naklon krivulje (b) se je gibal med 0,409 pri vrsti Thunnus thynnus in 4,343 pri vrsti Dentex macrophthalmus. Povprečna vrednost b je bila 3,088 (± SD = 0,898) in se ni bistveno razlikovala od 3 (t-test, p <0,05). Mediana koeficienta b je bila 3,062 in 50% vrednosti b se je gibalo med 2,941 in 3,190.
Ključne besede: razmerje med dolžino in težo, ribe, turška morja, pregled
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